NatureScot Research Report 1265 - Biological analyses of underwater video footage collected from around the Isle of Arran (Firth of Clyde) in 2015

Year of publication: 2021
Author: Dr Colin Moore

Cite as: Moore, C.G. 2021. Biological analyses of underwater video footage collected from around the Isle of Arran (Firth of Clyde) in 2015. NatureScot Research Report 1265.

The surveys were undertaken by NatureScot in July and September 2015, the latter in association with Marine Scotland Science. Most of the drop-down video runs were completed within a series of draft monitoring boxes.

Infaunal grab samples were also collected in the Arran monitoring boxes during the 2015 surveys. The results from the analyses of these samples were published in two discrete NatureScot reports in 2017 (Commissioned Report 945 and Commissioned Report 946) and have been related to the interpretation of the video sampling in the current report.

Keywords

benthos; biotope; PMF; MPA; protected feature; Annex I; habitat; video; Isle of Arran; Firth of Clyde

Background

The aim of this study was to improve knowledge of the occurrence and distribution of species and habitats of recognised conservation importance within and adjacent to the South Arran Marine Protected Area (MPA) through the analysis of seabed video and still photographic imagery collected during two surveys in 2015. The work also contributes to programmes of MPA monitoring and of refining knowledge of the distribution of Habitats Directive Annex I habitats within MPAs.

Main findings

The MPA Protected Feature (PF) ‘kelp and seaweed communities on sublittoral sediment’ was found to be widely distributed, particularly along the eastern coastline of Arran, generally in the form of dense Saccharina latissima and a red algal turf on maerl and stone gravel.
A single record of the ‘maerl or coarse shell gravel with burrowing sea cucumbers’ PF was located south of Holy Island.
Fairly extensive examples of the ‘maerl beds’ PF were observed at two locations. North-east of Pladda the bed was composed of low densities of live maerl, but in Lamlash Harbour overall live coverage was around 45%, although the maerl supported a relatively low epibiotic diversity for the habitat.
The ‘burrowed mud’ PF was extensively distributed around the Arran coastline over a depth range of 42 - 127 m. Seapen density was generally low. The megafaunal burrowing populations were low - moderately well-developed in shallower waters but were swelled by dense populations of Calocaris macandreae at depths of over 100 m. A single colony of the burrowed mud component species Funiculina quadrangularis was recorded.
In addition to the above PFs, which have also been designated as Priority Marine Features (PMFs), further PMFs observed include single individuals of the ocean quahog (Arctica islandica) at two locations, and a dense shoal of sandeels (Ammodytes sp.), although poor visibility precluded firm identification of the latter.
The ‘kelp and seaweed’, ‘maerl or coarse shell gravel’ and ‘maerl beds’ records are also categorised as Habitats Directive Annex I habitat sandbank features. Other sandbank feature records included three examples of megarippled maerl and stone gravel ascribed to the sandbank sub-type ‘gravelly and clean sands’. In addition, the sandbank sub-type ‘mixed sediments’ was well represented in the shallower draft MPA monitoring boxes.
The Annex I Reefs habitat was recorded in the form of bedrock and stony sub-types. Surveyed examples of bedrock reefs were sparse and of low diversity. Stony reefs were more widely recorded, mostly as scattered cobbles and boulders on muddy or mixed sediments, only occasionally constituting the dominant substrate.
Presumptive trawling disturbance in the form of shallow, parallel ridge and furrow scars was observed in the burrowed mud habitat of all monitoring boxes supporting the feature, apart from two, where the sampling intensity was relatively low.

Acknowledgements

Dr Graham Saunders is thanked for his contribution to QA of the project outputs. For the acquisition of imagery and associated survey data the author is grateful to the scientific staff and vessel crew of NatureScot and Marine Scotland Science, and to Ben James (NatureScot) for project management.

Introduction

This report contributes to a long series of underwater video analyses commissioned by SNH since 2009. The primary aim of these studies is to improve knowledge of the occurrence and distribution of species and habitats of recognised conservation importance through the analysis of seabed video and still photographic imagery collected during monitoring and research cruises. The current report aims to extend such knowledge for the South Arran Marine Protected Area (MPA) established in 2014, through the analysis of imagery collected during two surveys in 2015.

Drop-down video sampling was undertaken within and adjacent to the South Arran MPA (see Figure 1) in 2015 as part of the establishment of detailed, baseline monitoring of specified Protected Features (Table 1). This work was completed by NatureScot during two discrete surveys in July and September 2015, the latter in association with Marine Scotland Science. Most video runs were completed within a series of draft monitoring boxes.

Table 1. Protected Features of the South Arran MPA.

MPA protected features	Component biotope code/species name
Burrowed mud	SS.SMu.CFiMu.SpnMeg SS.SMu.CFiMu.MegMax Funiculina quadrangularis Pachycerianthus multiplicatus
Kelp and seaweed communities on sublittoral sediment	SS.SMp.KSwSS (excluding SS.SMp.KSwSS.Tra and SS.SMp.KSwSS.FilG)
Maerl beds	SS.SMp.Mrl
Maerl or coarse shell gravel with burrowing sea cucumbers	SS.SCS.CCS.Nmix
Ocean quahog aggregations	Arctica islandica
Seagrass beds	SS.SMp.SSgr.Zmar
Shallow tide-swept coarse sands with burrowing bivalves	SS.SCS.ICS.MoeVen

The conservation importance of features found in this report has been assessed with respect to a number of legislative drivers. These include the presence of Protected Features (PFs) and the presence of Scottish Priority Marine Features (PMFs) (SNH & JNCC, 2020). Cognisance has also been taken of other published importance measures. The current study also serves to contribute to a programme of refining the distribution of Habitats Directive Annex I habitats (European Commission, 2014) within Scottish protected areas.

Infaunal grab samples were also collected in the Arran monitoring boxes during the 2015 surveys. The results from the analyses of these samples have been published in two separate reports (Allen, 2017 and Axelsson et al., 2017) and have been related to the interpretation of the video sampling in the current report.

To aid the conservation objectives of the South Arran MPA, legislation to afford permanent protection to seabed habitats from physical disturbance by fishing came into effect in 2016 (i.e. after the surveys reported here) through the South Arran Marine Conservation Order 2015 (Scottish Government, 2015). This includes provisions to restrict the use of mobile, demersal fishing gear within the MPA and includes a ban on dredging and trawling within the whole of the MPA, with the exception of that by smaller trawlers (<120 tonnes) in deeper waters (Figure 1).

Map of the southern end of the Isle of Arran in the Clyde Sea showing the monitoring boxes and tracks of the drop-down video transects — Figure 1. Location of survey area around Arran, with inset showing wider geographical context.

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Also shown is the location of draft MPA monitoring boxes and video runs for the July *Seol Mara* and September *Alba na Mara* 2015 surveys, as well as prohibition zones for mobile, demersal fishing. Land mapping: © Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Methods

Survey details are provided in Table 2. Video images were obtained from dropdown video drifts, with the camera frame carrying a laser scaling system and the facility to take digital still photographs at intervals. For both surveys a high definition video camera captured HD imagery and also transmitted SD imagery to the surface vessel in real time. A video overlay on the SD footage displayed time and positional data, although the latter was unreliable for the Seol Mara survey. In fact vessel track log data were used for both surveys as this facilitated the subsequent production of high resolution, biotope-tagged track mapping. Trackpoint frequency was ca. 1 Hz for the Alba na Mara survey but only ca. 0.02 Hz for the Seol Mara survey, so for the latter track data frequency was increased to 1 Hz using linear interpolation. Depth data was provided on the video overlay in the case of the Alba na Mara survey. For the Seol Mara survey only the depths recorded at the start and end of the video runs were available. All depths were converted to depth below chart datum, employing TotalTide software (Admiralty, Taunton) to determine tidal rise at the most appropriate secondary port (Brodick Bay).

Table 2. Survey details
Organisation	Vessel	Location	Dates	No. sites	No. stills
SNH	MRV Seol Mara	East of Arran	14-20 July 2015	32	1064
SNH / MSS	MRV Alba na Mara	South of Arran	17-21 Sep 2015	25	1079

The video and still imagery was used to describe the nature of the seabed in terms of the physical structure and the species assemblages. Species present were, as far as possible, identified and quantified using the semi-quantitative MNCR SACFOR scale (Hiscock, 1996). Biotopes were allocated based on the physical and biological attributes (Connor et al., 2004). Runs traversing a sequence of habitats were split into corresponding segments, with the transition points recorded using the time, position and depth where the data were available. Video segments, as well as homogeneous, unsegmented runs, are regarded as video samples. Segmentation of runs was not practicable in the case of mosaics of biotopes, in which case all biotopes observed were simply listed. The presence of PFs, PMFs and Habitats Directive Annex I habitats was recorded for all video samples.

In the distributional maps of this report the positions of records have been plotted using either symbols marking the midpoint of the start and end of the sample, or using detailed vessel track data segmented into biotopes.

Results

Full details of the habitats, biotopes and species recorded along all video runs are provided in Annex 2, with location, temporal and depth data in Annex 1. The distribution of habitats, biotopes and species is summarised in the following sections for each of the monitoring boxes, as well as for adjacent regions where relevant. However, some broad distributional patterns can be discerned within the surveyed area as a whole (Figure 2) and these are outlined here. Within the Results section of the report PMF biotopes and species are highlighted using bold, italic text. Annex 3 provides an inventory of the biotopes recorded, together with illustrative photographs or video frame grabs and lists of their occurrence.

The great majority of biotope records fell within five broad categories and these tended to reflect a gradient of increasing depth. In the shallowest waters (6 - 12 m), which were sampled mostly off the east coast of Arran, the biota was dominated by fields of often dense Saccharina latissima (= Laminaria saccharina) and a red algal turf overlying sediments (SS.SMp.KSwSS), composed especially of maerl gravel and stone gravel (SS.SMp.KSwSS.LsacR.Gv). These algal-dominated habitats tended to give way to faunally-dominated mixed substrates over a depth range of mostly 17 - 35 m (SS.SMx.CMx). At many of these sites a substrate of predominantly silty, sandy gravel or gravelly sand supported dense Cerianthus lloydii (SS.SMx.CMx.ClloMx), which was often accompanied by frequent hydroids such as Nemertesia ramosa and N. antennina, as well as solitary ascidians, where significant quantities of shells and stones were also present (SS.SMx.CMx.ClloMx.Nem). Such mixed substrates sometimes were interrupted by patches of more homogeneous sediments, particularly in box D4 off the south of the island, but there was a tendency for them to be replaced by non-cohesive muddy sand (SS.SSa.CMuSa) and then cohesive muddy sand and sandy mud (SS.SMu.CSaMu) with increasing depth over a range of around 30 - 50 m. These cohesive sediments often supported sparse megafaunal burrows and sometimes Virgularia mirabilis, but from around 40 - 50 m depth gave way to burrowed mud habitats (SS.SMu.CFiMu.SpnMeg) which extended to 127 m. These were recorded widely around the island and were dominated by a burrowing fauna of Nephrops norvegicus and Calocaris macandreae, as well as generally low densities of V. mirabilis and sometimes Pennatula phosphorea. At the shallower sites (<100 m depth) the burrowing fauna was often poorly developed, but along the deeper runs within box T4 off the south-east of the island, dense populations of N. norvegicus and C. macandreae were recorded in soft mud, although seapens were sparse.

Map of the southern end of the Isle of Arran in the Clyde Sea showing the distribution of seabed habitats or biotopes observed along drop-down video monitoring transects — Figure 2. Distribution of biotope records around the Isle of Arran. Distribution within and adjacent to the monitoring boxes shown in greater detail in Figures 3 - 11. Also shown are prohibition zones for mobile, demersal fishing within the MPA.

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Land mapping: © Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Seol Mara July 2015 survey

As depth was only recorded at the start and end of video runs during this survey, indication of the depth range of the component biotope segments for those runs traversing more than one biotope is only available through reference to bathymetric chart data.

Box D3 (Figure 3)

There was a distinct west to east gradation in biotopes correlating with increasing depth in box D3. Inshore runs straddled the 10 m contour and traversed coarse sediment dominated by maerl and stone gravel supporting dense Saccharina latissima. This was accompanied by an algal turf dominated by Bonnemaisonia asparagoides, together with Desmarestia spp. and Chorda filum. Amongst the fauna there were high densities of Marthasterias glacialis and Ascidiella aspersa. The habitat has been ascribed principally to SS.SMp.KSwSS.LsacR.Gv, but also to SS.SMp.KSwSS.LsacR.Sa in sand-dominated patches. With increasing depth the substrate transitioned to silty sand and gravel supporting dense Cerianthus lloydii and Ascidiella aspersa (SS.SMx.CMx.ClloMx). This was recorded from around the 10 m contour to at least 23 m depth. Extensive areas of mixed sediment supporting a similar fauna were also present in deeper waters straddling the 30 m contour; however, scattered stones and shells provided a habitat for frequent hydroids, particularly Nemertesia ramosa and N. antennina (SS.SMx.CMx.ClloMx.Nem). Along run D3-5 this habitat formed a mosaic with boulders densely clothed with hydroids and ascidians, as well as Antedon species (CR.LCR.BrAs.AntAsH). Beyond the 30 m contour to 57 m depth, the maximum recorded within the box, there was a gradation from non-cohesive muddy sand to cohesive muddy sand and sandy mud such that precise habitat boundaries were difficult to determine. Evidence of the faunal composition was fairly similar over much of this depth range and included small holes, worm casts, emergent infaunal tubes and sparse Cerianthus lloydii. The harder, non-cohesive sediments have been ascribed to SS.SSa.CMuSa. The shallower, cohesive sediments along runs D3-3 and D3-5 also supported very sparse, small megafaunal burrows and so have been referred to SS.SMu.CSaMu. Nephrops norvegicus was common in cohesive muddy sand and sandy mud sediments in the deepest waters surveyed (46 - 57 m) along runs D3-3 and D3-6. The habitat represents a fairly poor example of the biotope SS.SMu.CFiMu.SpnMeg, with a low density of other megafaunal species and an absence of seapens.

The video-based interpretation of biotope distribution in box D3 is broadly similar to that identified from the infaunal grab data (Allen, 2017). Allen (2017) recorded an outer band of the sandy mud biotope SS.SMu.CSaMu.AfilMysAnit and an inner band of the shallow, mixed coarse sediment biotope SS.SMx.IMx. The latter corresponds to the zone of SS.SMp.KSwSS.LsacR.Gv recorded by video, but which would be difficult or impossible to identify from grab data alone.

The distribution of seabed habitats or biotopes observed along drop-down video monitoring transects and at point grab sampling stations in Arran monitoring box D3 — Figure 3. Distribution of biotope records along video runs and for grab samples in box D3.

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Land mapping: Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Box T1 (Figure 4)

Sediments in box T1 were muds spanning the depth range 71 - 95 m. The mud supported a megafaunal burrowing community of moderate density including Nephrops norvegicus, Calocaris macandreae and Lumpenus lampretaeformis and sparse seapens in the form of Virgularia mirabilis and Pennatula phosphorea (SS.SMu.CFiMu.SpnMeg). Along the central section of run T1-3 scattered cobbles and boulders supporting a hydroid turf formed a mosaic with the mud, in which only sparse small burrows were present (CR.LCR and SS.SMu.CFiMu).

Runs T1-7 and T1-8 were located around 500 m to the west of box T1 at a depth of 50 - 65 m. The predominant habitat type was sandy mud supporting a fairly light density of megafaunal burrows, although Nephrops norvegicus was common. Low densities of seapens were also present in the form of Virgularia mirabilis and a single example of Funiculina quadrangularis along run T1-8. In view of the fairly poorly-developed burrowing fauna and the sparsity of F. quadrangularis (only a single specimen being recorded during the July and September surveys), these muds have been referred to SS.SMu.CFiMu.SpnMeg. Along run T1-7 scattered, silted boulders and cobbles supported a hydroid turf (CR.LCR) amongst mud with few or no burrows (SS.SMu.CFiMu), with an area of silted, hydroid-dominated bedrock at the end of the run (CR.LCR).

The box was well covered by the grab survey in 2015, all samples being referred to the biotope SS.SMu.CFiMu (Allen, 2017). This is consistent with the current interpretation given the difficulty in identification of the presence of the characterising megafaunal constituents of SS.SMu.CFiMu.SpnMeg from grab data alone.

Box D5 and run LBN-1 (Figure 5)

The shallowest region surveyed within box D5 lay in the north-western part, where sediments dominated by maerl gravel were recorded from a depth of 7 m to around the 10 m contour. Living maerl was very sparse, the dead maerl material supporting a generally dense cover of Saccharina latissima and a patchy red algal turf including Bonnemaisonia asparagoides, as well as Desmarestia species and Chorda filum. Dominant faunal constituents included Cerianthus lloydii and Marthasterias glacialis, with Neopentadactyla mixta also being recorded (SS.SMp.KSwSS.LsacR.Gv). In this more sheltered region of the box from around the 10 m contour to a depth of 30 m mixed sediments of muddy sand and gravel predominated before giving way to muddy sand. The mixed sediments supported generally high numbers of Cerianthus lloydii, with stones and shells providing a substrate for dense Ascidiella aspersa, as well as for hydroids including Nemertesia ramosa and N. antennina (SS.SMx.CMx.ClloMx.Nem). The deeper muddy sand was probably transitional between SS.SSa.CMuSa and SS.SMu.CSaMu but has been ascribed to the latter in view of the recording of a single megafaunal burrow and occasional juvenile Virgularia mirabilis.

Runs in the south-eastern region of the box traversed deeper water, mostly from around 30 m to 52 m depth. There was a general depth-related gradient from mixed sediments through non-cohesive muddy sand to cohesive muddy sand or sandy mud in the deepest water. The mixed sediments supported a similar fauna to that of the same substrate type in the north-western region (SS.SMx.CMx.ClloMx.Nem), although Cerianthus lloydii and hydroid density diminished with depth, where a switch to the biotope SS.SMx.CMx was recognised. However, along run D5-2 scattered boulders supported dense hydroid turfs, together with Antedon species and a fairly rich ascidian fauna, resulting in the recognition of the additional biotope CR.LCR.BrAs.AntAsH. The mixed sediments graded into non-cohesive muddy sand which retained lower densities of some of the faunal constituents of the former substrate, such as hydroids, ascidians and Cerianthus lloydii (SS.SSa.CMuSa). This habitat formed a gradual transition into softer, muddy sand and sandy mud which had the cohesion to support very sparse megafaunal burrows. The infauna was also represented by small holes, worm casts and emergent tubes (SS.SMu.CSaMu). Runs D5-1 and D5-2 extended into the deepest water, where burrowed mud was recorded over a depth range of at least 46 - 52 m. Nephrops norvegicus burrows were common but the megafaunal burrowing community was otherwise poorly developed and no seapens were observed (SS.SMu.CFiMu.SpnMeg). This poor development of the biotope was similar to that observed over a similar depth range elsewhere off the east coast of Arran (box D3), but inferior to that recorded at similar depths off the south coast (boxes T2 and T3), where the sediment appeared softer. The burrowed mud habitat was interspersed by a small bedrock cliff and boulders along run D5-2. The silted rock supported occasional ascidians and a patchy hydroid turf (CR.LCR.BrAs).

Few of the grab samples taken during the survey were collected in the close vicinity of the video runs, but for those that were the interpretation of biotope identity was similar (Allen, 2017). However, as in the case of box D3 grab samples falling within the belt of SS.SMp.KSwSS.LsacR.Gv, Allen (2017) referred these shallow gravel sediments to mixed (SS.SMx.CMx) or coarse sediment (SS.SCS.ICS) biotopes in the absence of detailed algal data.

Run LBN-1 was located 400 m to the north-west of box D5 in an area lacking corresponding grab sample data. The run straddled the 10 m depth contour with a depth of 12 m recorded at both start and end. Half the run passed through a rich maerl bed with a width of around 250 m. Although rich in terms of live maerl density (overall around 45% cover), epibiotic diversity appeared relatively low for the habitat. The substrate of dense shells and maerl supported a short turf of filamentous red algae and frequent Saccharina latissima (Figure 14 A). The habitat has been assigned to the Phymatolithon calcareum biotope SS.SMp.Mrl.Pcal.R, although there is uncertainty regarding the identity of the dominant maerl species. The filamentous red algal turf and frequent S. latissima persisted for much of the rest of the video run but largely on a substrate of dense shells on silty sand (SS.SMp.KSwSS.LsacR). However, at the start of the run the habitat more closely resembled that recorded in the shallow, coarse sediments of box D5, with a substrate of maerl gravel supporting an algal flora dominated by filiform red and brown species, together with S. latissima (SS.SMp.KSwSS.LsacR.Gv).

The distribution of seabed habitats or biotopes observed along drop-down video monitoring transects and at point grab sampling stations within and adjacent to Arran monitoring box D5 — Figure 5. Distribution of biotope records along video runs and for grab samples in box D5 and adjacent waters.

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Land mapping: Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Box D1 and T5 runs (Figure 6)

The northern, shallower region of box D1 was floored by maerl gravel dominated sediments supporting very sparse live maerl thalli (<1%) and extending shorewards to a depth of 7m. Saccharina latissima was generally frequent - abundant, accompanied by a patchy filiform red algal turf and several species of Desmarestia. Faunal dominants included often dense Cerianthus lloydii and asteroid echinoderms, particularly Marthasterias glacialis (SS.SMp.KSwSS.LsacR.Gv). At a depth of around 10 m along the most exposed run (D1-4) the dead maerl was formed into megaripples and the algal component was much reduced. This area has been ascribed to SS.SCS.ICS, although the presence of abundant C. lloydii is atypical of the biotope. Between the 10 and 20 m depth contours within the box the habitat transitioned to mixed sediments of silty sandy gravel and silty gravelly sand supporting dense C. lloydii (SS.SMx.CMx.ClloMx), extending to a depth of 37 m. However at D1-4, following a small patch of dense Ophiocomina nigra (SS.SMx.CMx.OphMx), the anemones were accompanied by frequent Nemertesia species for much of the run (SS.SMx.CMx.ClloMx.Nem). In the silty, sandy gravel along D1-2 the community was similar to that observed in the SS.SMx.CMx.ClloMx grounds. However, it has been referred to the burrowing holothurian biotope SS.SCS.CCS.Nmix due to the recorded presence of a single individual of Neopentadactyla mixta. The density of this species is difficult to determine due to its sediment withdrawal behaviour and so this biotope may be far more widespread. Run D1-6 was located to the south of the other runs, mostly in deeper water extending to 43 m depth. The sediment was a cohesive muddy sand interrupted by areas of slightly mixed muddy sand with gravel and scattered pebbles and shells, although there were broad transitional zones. The softer, cohesive sediment areas supported very sparse small burrows, as well as Turritella communis and Ophiura ophiura, with stones and shells providing a substrate for Nemertesia species and Omalosecosa ramulosa (SS.SMu.CSaMu). The harder, more mixed substrate areas supported a similar fauna, except for a lack of burrows (SS.SMx.CMx).

The interpretation of biotope distribution in box D1 based on imagery is broadly in line with that derived from the contemporaneous grab survey (Allen, 2017). Allen (2017) recorded SS.SMx.CMx in the vicinity of areas identified as SS.SMx.CMx biotopes based on video analysis. SS.CCS.ICS was also recorded by both surveys in a similar location and both surveys identified the presence of SS.SMu.CSaMu biotopes in the deeper, southern region of the box. As in the case of boxes D3 and D5, one grab sample was allocated to SS.SMx.IMx within a belt of ground identified as SS.SMp.KSwSS.LsacR.Gv from the video evidence.

Three T5 runs were located on sandy mud 300 - 1300 m to the east of box D1 at a depth of 70 - 84 m. The deepest run T5-2 traversed a fairly lightly burrowed sediment, although Nephrops norvegicus was common and Calocaris macandreae increased in density along the run, becoming common as the sediment softened with depth. Occasional Virgularia mirabilis was also present (SS.SMu.CFiMu.SpnMeg). A similar fauna was recorded for most of the distance along the shallower runs, T5-1 and T5-3, although the megafaunal burrowing community was relatively poorly developed (SS.SMu.CFiMu.SpnMeg) and was virtually absent at the end of run T5-3, with a gradual transition to slightly harder sediment supporting Ophiura ophiura, Arctica islandica and frequent V. mirabilis (SS.SMu.CSaMu).

Possible trawl scars were observed in the sediment over a small area along run T5-1.

No grab samples were collected from the vicinity of the T5 runs (Allen, 2017).

Alba na Mara September 2015 survey

Box T4 (Figure 7)

All runs in this box traversed significantly greater depths than the boxes described elsewhere (114 - 127 m) and were over soft muds supporting a dense megafaunal burrowing community dominated by Nephrops norvegicus (common) and Calocaris macandreae (SS.SMu.CFiMu.SpnMeg). Density of the latter was difficult to discern due to overlapping clusters of burrows but was within the range common - abundant, attaining the latter density category at least locally. Virgularia mirabilis was present at low density. Possible parallel trawl disturbance marks were recorded along three of the five runs (Figure 15 C). Axelsson et al. (2017) recorded a wide range of mud content in grab samples from this box (17 - 81%) but ascribed them all to SS.SMu.CSaMu.

Box D6 (Figure 8)

Video runs in box D6 spanned a wide depth range from 14 to 44 m. The innermost video run, D6-V03, was markedly shallower (14 - 22 m) than the others and had the most distinctive habitat composition. Most of the run traversed coarse sediments with a high maerl gravel content, in places formed into megaripples. The sediments in the central third of the run over a distance of around 650 m supported frequent live maerl thalli with around 10% cover. The algal flora was otherwise poorly developed and included occasional filamentous and filiform reds, Desmarestia aculeata and frequent, though probably largely drift, Saccharina latissima (SS.SMp.Mrl) (Figure 14 B). The visible fauna was dominated by echinoderms, as well as Ascidiella aspersa, which was abundant in patches. On each side of the maerl bed, the habitat transitioned into megarippled coarse sediments with sparse live maerl (<1%) and other algae, but which supported a similar fauna to that of the maerl bed (SS.SCS.CCS). Towards the northern end of the run scattered pebbles, cobbles and boulders on gravelly sand provided a substrate for a patchy red algal turf, dense Cerianthus lloydii and superabundant aggregations of Ascidiella aspersa (SS.SMp.KSwSS).

In deeper water the dominant habitat recorded in box D6 was a mixed, gravelly sand sediment supporting very dense populations of Cerianthus lloydii. This has been only tentatively ascribed to the biotope SS.SMx.CMx.ClloMx in view of the atypically low mud content of the sediment. Scattered shells, pebbles, cobbles and boulders provided a substrate for sparse hydroids, Flustra foliacea and red algae. Along run D6-V04 the community was supplemented by patches of dense Ophiocomina nigra (SS.SMx.CMx.OphMx). In the deepest water from 37 - 44 m the mixed gravelly sand sediments acquired a significant silt content and the dense Cerianthus lloydii populations were lost (SS.SMx.CMx). Areas of slightly muddy fine sand supporting Turritella communis were also recorded here (SS.SSa.CMuSa). In spite of their shallow depth (34 - 52 m), Axelsson et al. (2017) ascribed all grab samples collected within the box to the offshore sand biotope, SS.SSa.OSa.OfusAfil.

The distribution of seabed habitats or biotopes observed along drop-down video monitoring transects and at point grab sampling stations in monitoring box D6 — Figure 8. Distribution of biotope records along video runs and for grab samples in box D6 and adjacent waters.

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Land mapping: Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Box T2 (Figure 9)

Depths in box T2 spanned the range 44 to 56 m with a transition from cohesive muddy sand to soft mud occurring at around the 50 m contour. Beyond the 50 m contour the mud supported a similar megafaunal burrowing community to that of box T3, with Nephrops norvegicus (common) and Calocaris macandreae accompanied by low densities of Pennatula phosphorea and Virgularia mirabilis (SS.SMu.CFiMu.SpnMeg). In predominantly shallower waters the muddy sand along runsT2-V04 and T2-V05 supported sparse burrows and Turritella communis and has been ascribed to SS.SMu.CSaMu. Possible trawl disturbance marks in the form of parallel ridges and furrows were observed at several points along run T2-V06 (Figure 15 A, B). Despite the high mud content (up to 78%) found in grab samples in the southern region of the box, Axelsson et al. (2017) have referred the samples to the muddy sand biotope SS.SSa.CMuSa.AalbNuc. However, biotope determination by Axelsson et al. (2017) of grab samples from the shallower, northern region of the box (SS.SMu.CSaMu.AfilMysAnit) is consistent with the current video interpretation.

Box D4 (Figure 10)

Video runs in box D4 traversed a depth range of 24 to 36 m. Mixed sediments of silty sand and gravel, often with significant quantities of pebbles, were widely distributed throughout the area with evidence of an infaunal component often in the form of small mounds, holes and emergent tubes, while stones and shells supported encrusting pink coralline algae, hydroids and Omalosecosa ramulosa. The motile fauna was dominated by echinoderms, particularly Marthasterias glacialis, Asterias rubens and Porania pulvillus (SS.SMx.CMx). Silted cobbles and boulders were scattered throughout much of the area but locally formed dense patches, sometimes as a mosaic with mixed sediments. These larger stones supported a similar biota to the smaller stones but with a denser hydroid cover, as well as ascidians such as Ascidia mentula and A. virginea (CR.LCR.BrAs). In deeper water (29 - 32 m) the stones also supported low numbers of Antedon spp. (CR.LCR.BrAs.AntAsH). Areas of more homogeneous sediments were also recorded along most of the video runs in the box. In shallower waters (29 - 33 m) along runs D4-V01 and D4-V04 these were in the form of slightly muddy fine sand punctured by small holes and with small mounds and sparse emergent infaunal tubes (SS.SSa.CMuSa). In deeper waters (33 - 36 m) along runs D4-V03 and D4-V05 cohesive muddy sand supported Turritella communis, Ophiura ophiura and sparse megafaunal burrows (SS.SMu.CSaMu). It should be noted that grab samples collected contemporaneously with the video survey throughout box D4, with many in close proximity to the video runs and over a similar depth range (29 - 40 m), have all been ascribed to offshore sand biotopes (SS.SSa.OSa, SS.SSa.OSa.OfusAfil) by Axelsson et al. (2017). The video evidence suggests a different interpretation, particularly in view of the filamentous and foliose red algae recorded at some sites and megafaunal burrows at others.

Box T3 (Figure 11)

Video analysis within box T3 suggested that the predominant substrates were soft muds and sandy muds, although particle size analysis of grab samples recorded chiefly muddy sand and some slightly sandy mud (Axelsson et al., 2017). These cohesive muddy sediments in the main supported a moderate density of megafaunal, crustacean burrowers, with Nephrops norvegicus (common), accompanied by Calocaris macandreae and Goneplax rhomboides. Seapens were sparse in the form of Pennatula phosphorea and Virgularia mirabilis. The three deeper video runs (46 - 54 m depth) are referable to SS.SMu.CFiMu.SpnMeg on the basis of video evidence, although corresponding grab samples have been ascribed to SS.SSa.CMuSa biotopes (Axelsson et al., 2017). Megafaunal burrow density was slightly reduced in the sandier muddy sediments along the shallowest video run (T3-V03) at 42 - 48 m depth. Nephrops norvegicus was still common but accompanied by high numbers of Turritella communis. The habitat was transitional between SS.SMu.CFiMu.SpnMeg and SS.SMu.CSaMu but has been ascribed to the former. Mixed substrate patches were also recorded along this run composed of muddy sediments with scattered gravel, pebbles, cobbles and boulders supporting a hydroid turf (SS.SMx.CMx). Possible evidence of trawl ground gear disturbance of the seabed was present along two of the runs (Figure 15 D).

Discussion

This section considers the conservation importance of the species and habitats encountered during the surveys, while also providing a summary appraisal of the distribution of PMFs and, where relevant, MPA protected features and Habitats Directive Annex I habitats. The conservation importance of species and habitats and their occurrence in the Arran surveys are summarised in Table 3, with the occurrence of Annex I habitats given in Table 4. The distribution of Annex I habitat records is shown in Figure 12 and those of PFs and PMFs in Figure 13. Annex 2 includes details of all Annex I, PF and PMF records.

Around half the biotopes in Table 3 fall within broad habitat types included in the Scottish Biodiversity List (Scottish Government, 2013) but are not recognised by other indicators of conservation importance. They are generally of wide occurrence.

Table 3. Species and biotopes recorded during the Arran surveys of recognised conservation importance and their frequency of occurrence in video samples. Importance indicators are SBL = Scottish Biodiversity List of Habitats and Species, Osp = OSPAR List of Threatened and/or Declining Species and Habitats, PMF = Priority Marine Feature, PF = Protected Feature. Abbreviations used in the tables of this report for PMFs and PFs are provided in Annex 4.

Table 3.
Biotope/species	SBL	Osp	PMF	PF	# records
SS.SCS.ICS	Yes	No	No	No	1
SS.SCS.CCS	Yes	No	No	No	3
SS.SCS.CCS.Nmix	Yes	No	Yes	Yes	1
SS.SSa.CMuSa	Yes	No	No	No	19
SS.SMu.CSaMu	Yes	No	No	No	20
SS.SMu.CFiMu	Yes	No	No	No	3
SS.SMu.CFiMu.SpnMeg	Yes	Yes	Yes	Yes	35
SS.SMx.CMx	Yes	No	No	No	35
SS.SMx.CMx.ClloMx	Yes	No	No	No	17
SS.SMx.CMx.ClloMx.Nem	Yes	No	No	No	11
SS.SMx.CMx.OphMx	Yes	No	No	No	7
SS.SMp.Mrl	Yes	Yes	Yes	Yes	1
SS.SMp.Mrl.Pcal.R	Yes	Yes	Yes	Yes	2
SS.SMp.KSwSS	No	No	Yes	Yes	1
SS.SMp.KSwSS.LsacR	No	No	Yes	Yes	3
SS.SMp.KSwSS.LsacR.Gv	No	No	Yes	Yes	13
SS.SMp.KSwSS.LsacR.Sa	No	No	Yes	Yes	1
Arctica islandica	No	No	Yes	No	2
Funiculina quadrangularis	No	No	Yes	Yes	1
Ammodytes spp.?	No	No	Yes	No	1

Table 4. Frequency of occurrence of Annex I habitats recorded during the Arran surveys. Abbreviations used in the tables of this report for habitats are provided in Annex 4.

Habitat	Sub-type/feature	# records
Reefs	Bedrock	2
Reefs	Stony	17
Subtidal sandbanks	Gravelly & clean sands	4
Subtidal sandbanks	Mixed sediments	17
Subtidal sandbanks	Maerl beds	3
Subtidal sandbanks	Kelp and seaweed	17

Annex I habitats were represented by reefs and subtidal sandbanks (Figure 12, Table 4). The reefs were of low quality. Bedrock reefs consisted of sparse, small outcrops that were silted and supported a low diversity community. Stony reefs were more widely recorded, mostly as scattered cobbles and boulders on muddy or mixed sediments, only occasionally constituting the dominant substrate.

Sandbank habitats were observed within or in the vicinity of four of the monitoring boxes, where depths of less than 24 m were recorded. Mixed sediments belonging to the SS.SMx.CMx biotope complex were widespread. Three of the gravelly and clean sands Annex 1 habitat records consisted of megarippled maerl and stone gravel supporting dense Cerianthus lloydii (SS.SCS.ICS, SS.SCS.CCS). All the remaining sandbank habitat records were also both Priority Marine Features and MPA Protected Features. These included a single record of the ‘maerl or coarse shell gravel with burrowing sea cucumbers’ PMF (SS.SCS.CCS.Nmix). This was in a location, south of Holy Island, where the habitat has been historically widely recorded according to the GeMS database (v8, i24; SNH, 2020).

The geographical pattern of Annex I sandbank records reflects the greater intensity of shallow water sampling along the eastern coastline in comparison to the southern coastline. The ‘kelp and seaweed communities on sublittoral sediment’ PMF was widely recorded along the east coast, generally in the form of dense Saccharina latissima and a red algal turf on maerl and stone gravel (SS.SMp.KSwSS.LsacRGv).

The maerl bed PMF was recorded at two locations: Lamlash Harbour (two records briefly interrupted by a patch of low live maerl density along run LBN-1), and north-east of Pladda (just north of box D6). The video evidence suggests both beds were fairly extensive with distances of around 250 m and 650 m respectively being covered by the traversing camera. The Pladda bed was poorly developed with only around 10% live maerl cover and a fairly sparse epibiotic macroalgal flora (SS.SMp.Mrl) (Figure 14 A). The Lamlash Harbour site exhibited a much richer live maerl content (around 45% cover), which supported a dense filamentous algal turf but a relatively low epibiotic diversity for the habitat type (SS.SMp.Mrl.Pcal.R) (Figure 14 B).

A further habitat of conservation importance, burrowed mud (SS.SMu.CFiMu.SpnMeg), was recorded. This is not on the Annex I list, but is both a PF and PMF. This was extensively distributed around the Arran coastline over a depth range of 42 - 127 m. In general the seapen populations were poorly developed but the density of Nephrops norvegicus was typical for the habitat (common). Overall megafaunal burrow density was low to moderate in shallower waters, but was swelled by dense populations of Calocaris macandreae (common - abundant) at depths of over 100 m in box T4. A single colony of the seapen Funiculina quadrangularis (a burrowed mud PF and PMF component species) was observed in box T1.

Map of the southern end of the Isle of Arran in the Clyde Sea showing the presence and distribution of seabed communities correlating with European Commission Habitats Directive Annex I habitats — Figure 12. Distribution of Habitats Directive Annex I habitat records around the Isle of Arran, as well as prohibition zones for mobile, demersal fishing within the MPA.

Click for a full description

Land mapping: © Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Map of the southern end of the Isle of Arran in the Clyde Sea showing the presence and distribution of seabed communities correlating with protected features of the South Arran Marine Protected Area — Figure 13. Distribution of PF and PMF records around the Isle of Arran, as well as prohibition zones for mobile, demersal fishing within the MPA.

Click for a full description

Land mapping: © Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

Other PMF species recorded included single individuals of Arctica islandica in sandy mud at two locations along run T5-3 and a dense shoal of possibly Ammodytes sp. at the start of run D3-4 over mixed coarse sediment. Poor visibility precluded firm identification.

Maerl bed habitats — Figure 14. Photographic illustration of the two maerl beds recorded: A, Lamlash Harbour (run LBN-1); B, off Pladda (run D6-V03).

Presumptive trawling disturbance in the form of shallow, parallel scars (Figure 15) was observed at eight locations (Figure 16). All disturbance records were on burrowed mud (SS.SMu.CFiMu.SpnMeg) and included all monitoring boxes where the habitat was present, apart from boxes D3 and D5, where sampling intensity of the habitat was relatively low. Visibility of soft muddy habitats was generally fairly poor and this will have influenced the extent to which evidence of fishing activity will have been identified and has militated against assessment of the impact of the disturbance on the benthic community.

A series of four slightly blurred images captured from the drop-down video footage of muddy seabed habitats — Figure 15. Examples of presumptive trawl scars along video runs through areas of burrowed mud. Runs: A, T2-V06; B, T2-V06; C, T4; D, T3-V01. Red laser dots 6.4 cm apart, green laser lines 50 cm apart.

Map of the southern end of the Isle of Arran in the Clyde Sea showing the distribution of scars or furrows — Figure 16. Observations of presumptive trawling scars in relation to the distribution of burrowed mud records. Also shown are prohibition zones for mobile, demersal fishing within the MPA (introduced after the survey work was completed).

Click for a full description

Land mapping: © Crown copyright [and database rights] 2021 OS 100017908. Bathymetry: © Crown copyright 2021. All rights reserved. Licence No. EK001-20140401. Not to be used for navigation.

References

Allen, J.H. 2017. Infaunal and PSA analyses of benthic samples collected from around the Isle of Arran, Loch Fyne and Orkney in July and August 2015. Scottish Natural Heritage Commissioned Report No. 945.

Axelsson, M., O’Dell, J. & Dewey, S. 2017. Infaunal and PSA analyses of benthic samples collected from South Arran MPA, Lochs Duich, Long and Alsh MPA and Southern Trench MPA proposal. Scottish Natural Heritage Commissioned Report No. 946.

Connor, D.W., Allen, J.H., Golding, N., Howell, K.L., Lieberknecht, L.M., Northen, K.O. & Reker, J.B. 2004. The National Marine Habitat Classification for Britain and Ireland. Version 04.05. Peterborough: Joint Nature Conservation Committee.

European Commission, 2013. Interpretation Manual of European Union Habitats. European Commission DG Environment.

Hiscock, K. (ed). 1996. Marine Nature Conservation Review: Rationale and Methods. Peterborough: Joint Nature Conservation Committee.

Scottish Government, 2013. Scottish Biodiversity List. Scottish Government.

Scottish Government, 2015. Scottish Statutory Instruments 2015 No. 437. The South Arran Marine Conservation Order 2015. Scottish Government.

SNH, 2020. GeMS - Scottish Priority Marine Features (PMF). Scottish Natural Heritage.

SNH & JNCC, 2020. Priority Marine Features in Scotland's seas. SNH & JNCC.

Annex 1: Positional, temporal and depth details of video sequences recorded during the surveys

Where video runs at a site have been divided into segments of different habitats, the segments are coded Sample.x where x is the segment number. The Seol Mara survey is dated July 2015, the Alba na Mara survey September 2015.

Sample	Date	Video filename (HD)	Time code start HD	Time code end HD	Time start (UT)	Time end (UT)	Start latitude	Start longitude	End latitude	End longitude	Depth CD start (m)	Depth CD end (m)
T1-1	14/07/2015	Arran_T1-1_2015_07_14.mts	00:06:44	00:43:38	08:39:52	09:16:46	55.567958	-5.052102	55.564322	-5.065324	82.1	69.9
T1-2	14/07/2015	Arran_T1-2_2015_07_14.mts	00:11:02	00:31:16	09:45:53	10:06:07	55.577562	-5.060646	55.575613	-5.065966	94.8	92.7
T1-3.01	14/07/2015	Arran_T1-3_2015_07_14.mts	00:07:22	00:21:38	10:30:25	10:44:41	55.568143	-5.072692	55.570035	-5.068290	71.6	-
T1-3.02	14/07/2015	Arran_T1-3_2015_07_14.mts	00:21:38	00:22:22	10:44:41	10:45:25	55.570035	-5.068290	55.570123	-5.068010	-	-
T1-3.03	14/07/2015	Arran_T1-3_2015_07_14.mts	00:22:22	00:37:28	10:45:25	11:00:31	55.570123	-5.068010	55.571814	-5.063406	-	81.6
T1-4	14/07/2015	Arran_T1-4_2015_07_14.mts	00:06:58	00:32:06	11:24:11	11:49:19	55.566722	-5.068030	55.569605	-5.062475	70.6	75.7
T1-5	14/07/2015	Arran_T1-5_2015_07_14.mts	00:08:54	00:37:36	12:15:38	12:44:20	55.578992	-5.078834	55.580878	-5.070399	84.8	80.0
D5-1.01	14/07/2015	Arran_D5-1_2015_07_14.mts	00:05:27	00:17:36	13:23:51	13:36:00	55.531615	-5.061732	55.533417	-5.061746	52.4	-
D5-1.02	14/07/2015	Arran_D5-1_2015_07_14.mts	00:17:36	00:20:50	13:36:00	13:39:14	55.533417	-5.061746	55.533992	-5.061791	-	-
D5-1.03	14/07/2015	Arran_D5-1_2015_07_14.mts	00:20:50	00:24:38	13:39:14	13:43:02	55.533992	-5.061791	55.534809	-5.061670	-	-
D5-1.04	14/07/2015	Arran_D5-1_2015_07_14.mts	00:24:38	00:35:50	13:43:02	13:54:14	55.534809	-5.061670	55.537001	-5.061670	-	29.7
D5-2.01	14/07/2015	Arran_D5-2_2015_07_14.mts	00:05:20	00:10:39	14:06:49	14:12:08	55.532203	-5.065537	55.533239	-5.065514	46.8	-
D5-2.02	14/07/2015	Arran_D5-2_2015_07_14.mts	00:10:39	00:12:50	14:12:08	14:14:19	55.533239	-5.065514	55.533632	-5.065593	-	-
D5-2.03	14/07/2015	Arran_D5-2_2015_07_14.mts	00:12:50	00:21:32	14:14:19	14:23:01	55.533632	-5.065593	55.535459	-5.066026	-	-
D5-2.04	14/07/2015	Arran_D5-2_2015_07_14.mts	00:21:32	00:25:20	14:23:01	14:26:49	55.535459	-5.066026	55.536276	-5.066217	-	-
D5-2.05	14/07/2015	Arran_D5-2_2015_07_14.mts	00:25:20	00:26:43	14:26:49	14:28:12	55.536276	-5.066217	55.536518	-5.066212	-	-
D5-2.06	14/07/2015	Arran_D5-2_2015_07_14.mts	00:26:43	00:27:54	14:28:12	14:29:23	55.536518	-5.066212	55.536751	-5.066210	-	-
D5-2.07	14/07/2015	Arran_D5-2_2015_07_14.mts	00:27:54	00:31:58	14:29:23	14:33:27	55.536751	-5.066210	55.537546	-5.066317	-	-
D5-2.08	14/07/2015	Arran_D5-2_2015_07_14.mts	00:31:58	00:35:54	14:33:27	14:37:23	55.537546	-5.066317	55.538266	-5.066586	-	-
D5-2.09	14/07/2015	Arran_D5-2_2015_07_14.mts	00:35:54	00:37:03	14:37:23	14:38:32	55.538266	-5.066586	55.538533	-5.066609	-	-
D5-2.10	14/07/2015	Arran_D5-2_2015_07_14.mts	00:37:03	00:38:14	14:38:32	14:39:43	55.538533	-5.066609	55.538754	-5.066666	-	45.9
D5-3.01	14/07/2015	Arran_D5-3_2015_07_14.mts	00:02:54	00:15:58	14:53:19	15:06:23	55.533383	-5.069708	55.535736	-5.070053	19.6	-
D5-3.02	14/07/2015	Arran_D5-3_2015_07_14.mts	00:15:58	00:17:22	15:06:23	15:07:47	55.535736	-5.070053	55.535996	-5.070241	-	-
D5-3.03	14/07/2015	Arran_D5-3_2015_07_14.mts	00:17:22	00:18:46	15:07:47	15:09:11	55.535996	-5.070241	55.536342	-5.070116	-	-
D5-3.04	14/07/2015	Arran_D5-3_2015_07_14.mts	00:18:46	00:22:34	15:09:11	15:12:59	55.536342	-5.070116	55.537282	-5.070096	-	39.5
D5-4.01	14/07/2015	Arran_D5-4_2015_07_14.part 1.mts	00:02:34	00:04:34	15:22:45	15:24:44	55.536961	-5.074737	55.537019	-5.074658	8.6	-
D5-4.02	14/07/2015	Arran_D5-4_2015_07_14.part 1.mts	00:04:34	00:16:31	15:24:44	15:36:41	55.537019	-5.074658	55.539272	-5.076059	-	-
D5-4.03	14/07/2015	Arran_D5-4_2015_07_14.part 2.mts	00:01:27	00:03:40	15:41:52	15:44:05	55.539221	-5.075736	55.539621	-5.075815	-	-
D5-4.04	14/07/2015	Arran_D5-4_2015_07_14.part 2.mts	00:03:40	00:10:33	15:44:05	15:50:58	55.539621	-5.075815	55.541085	-5.076193	-	29.7
D5-5.01	14/07/2015	Arran_D5-5_2015_07_14.mts	00:02:08	00:13:26	16:01:14	16:12:32	55.538513	-5.081517	55.539430	-5.077197	8.5	-
D5-5.02	14/07/2015	Arran_D5-5_2015_07_14.mts	00:13:26	00:18:25	16:12:32	16:17:31	55.539430	-5.077197	55.539863	-5.075890	-	-
D5-5.03	14/07/2015	Arran_D5-5_2015_07_14.mts	00:18:25	00:19:36	16:17:31	16:18:42	55.539863	-5.075890	55.539930	-5.075402	-	30.5
D5-6.01	14/07/2015	Arran_D5-6_2015_07_14.mts	00:01:20	00:15:21	16:29:16	16:43:17	55.537956	-5.077253	55.539827	-5.080934	6.5	-
D5-6.02	14/07/2015	Arran_D5-6_2015_07_14.mts	00:15:21	00:19:03	16:43:17	16:46:59	55.539827	-5.080934	55.539837	-5.082400	-	19.8
D1-1.01	16/07/2015	Arran_D1-1_2015_07_16.mts	00:03:54	00:10:42	08:16:09	08:22:57	55.510829	-5.070087	55.509568	-5.069165	8.1	-
D1-1.02	16/07/2015	Arran_D1-1_2015_07_16.mts	00:10:42	00:22:59	08:22:57	08:35:14	55.509571	-5.069166	55.506814	-5.067285	-	-
D1-1.03	16/07/2015	Arran_D1-1_2015_07_16.mts	00:22:59	00:23:28	08:35:14	08:35:43	55.506814	-5.067285	55.506747	-5.067163	-	-
D1-1.04	16/07/2015	Arran_D1-1_2015_07_16.mts	00:23:28	00:23:58	08:35:43	08:36:12	55.506747	-5.067163	55.506679	-5.067043	-	17.9
D1-2.01	16/07/2015	Arran_D1-2_2015_07_16 part 1.mts	00:01:38	00:09:48	08:44:20	08:52:30	55.510761	-5.065732	55.508834	-5.064286	6.8	-
D1-2.02	16/07/2015	Arran_D1-2_2015_07_16 part 2.mts	00:01:19	00:09:18	08:56:03	09:04:03	55.508479	-5.064100	55.506767	-5.062527	-	-
D1-2.03	16/07/2015	Arran_D1-2_2015_07_16 part 2.mts	00:09:18	00:14:53	09:04:03	09:09:38	55.506767	-5.062527	55.505459	-5.061584	-	-
D1-2.04	16/07/2015	Arran_D1-2_2015_07_16 part 2.mts	00:14:53	00:19:34	09:09:38	09:14:19	55.505459	-5.061584	55.504399	-5.060998	-	36.6
D1-3.01	16/07/2015	Arran_D1-3_2015_07_16.mts	00:03:03	00:07:25	09:30:45	09:35:07	55.511276	-5.061744	55.510167	-5.060540	8.5	-
D1-3.02	16/07/2015	Arran_D1-3_2015_07_16.mts	00:07:25	00:22:25	09:35:07	09:50:06	55.510171	-5.060544	55.506332	-5.058090	-	28.3
D1-4.01	16/07/2015	Arran_D1-4_2015_07_16.mts	00:02:35	00:04:52	10:00:21	10:02:37	55.512912	-5.058515	55.512527	-5.057753	10.3	-
D1-4.02	16/07/2015	Arran_D1-4_2015_07_16.mts	00:04:52	00:06:50	10:02:37	10:04:35	55.512527	-5.057753	55.512005	-5.057325	-	-
D1-4.03	16/07/2015	Arran_D1-4_2015_07_16.mts	00:06:50	00:07:23	10:04:35	10:05:07	55.512005	-5.057325	55.511859	-5.057207	-	-
D1-4.04	16/07/2015	Arran_D1-4_2015_07_16.mts	00:07:23	00:20:23	10:05:07	10:18:10	55.511859	-5.057207	55.508622	-5.055499	-	35.1
D1-5.01	16/07/2015	Arran_D1-5_2015_07_16.mts	00:04:41	00:23:18	10:39:26	10:58:03	55.510798	-5.061011	55.509529	-5.067746	10.0	-
D1-5.02	16/07/2015	Arran_D1-5_2015_07_16.mts	00:23:18	00:29:36	10:58:03	11:04:21	55.509529	-5.067740	55.509212	-5.070084	-	26.8
D1-6.01	16/07/2015	Arran_D1-6_2015_07_16.mts	00:03:49	00:16:01	11:20:29	11:32:41	55.502494	-5.067292	55.503070	-5.062724	27.7	-
D1-6.02	16/07/2015	Arran_D1-6_2015_07_16.mts	00:16:01	00:26:09	11:32:41	11:42:49	55.503070	-5.062724	55.503796	-5.058273	-	-
D1-6.03	16/07/2015	Arran_D1-6_2015_07_16.mts	00:26:09	00:30:30	11:42:49	11:47:10	55.503796	-5.058273	55.504143	-5.056426	-	-
D1-6.04	16/07/2015	Arran_D1-6_2015_07_16.mts	00:30:30	00:33:31	11:47:10	11:50:11	55.504143	-5.056426	55.504460	-5.055233	-	42.6
T1-6	16/07/2015	Arran_T1-6_2015_07_16.mts	00:06:55	00:21:53	12:44:50	12:59:48	55.574364	-5.075060	55.575451	-5.069990	81.5	87.5
T1-7.01	16/07/2015	Arran_T1-7_2015_07_16.mts	00:04:53	00:05:05	13:23:42	13:23:54	55.575561	-5.090293	55.575502	-5.090267	51.1	-
T1-7.02	16/07/2015	Arran_T1-7_2015_07_16.mts	00:05:05	00:28:19	13:23:54	13:47:08	55.575502	-5.090267	55.570551	-5.085446	-	-
T1-7.03	16/07/2015	Arran_T1-7_2015_07_16.mts	00:28:19	00:28:47	13:47:08	13:47:36	55.570551	-5.085446	55.570432	-5.085341	-	-
T1-7.04	16/07/2015	Arran_T1-7_2015_07_16.mts	00:28:47	00:31:40	13:47:36	13:50:29	55.570432	-5.085341	55.569820	-5.084835	-	-
T1-7.05	16/07/2015	Arran_T1-7_2015_07_16.mts	00:31:40	00:31:44	13:50:29	13:50:32	55.569820	-5.084835	55.569807	-5.084829	-	49.9
T1-8	16/07/2015	Arran_T1-8_2015_07_16.mts	00:05:27	00:48:11	14:06:15	14:48:59	55.573983	-5.081968	55.566367	-5.076865	65.3	58.6
T1-9	16/07/2015	Arran_T1-9_2015_07_16.mts	00:07:18	00:43:01	15:12:00	15:47:43	55.577801	-5.063008	55.570013	-5.056421	94.3	81.0
LBN-1.01	18/07/2015	Arran_LBN_2015_07_18.mts	00:03:11	00:08:11	13:08:57	13:13:57	55.545330	-5.085503	55.545771	-5.086368	11.6	-
LBN-1.02	18/07/2015	Arran_LBN_2015_07_18.mts	00:08:11	00:09:38	13:13:57	13:15:24	55.545771	-5.086368	55.545775	-5.086746	-	-
LBN-1.03	18/07/2015	Arran_LBN_2015_07_18.mts	00:09:38	00:19:22	13:15:24	13:25:08	55.545775	-5.086746	55.546140	-5.090360	-	-
LBN-1.04	18/07/2015	Arran_LBN_2015_07_18.mts	00:19:22	00:20:42	13:25:08	13:26:28	55.546140	-5.090360	55.546198	-5.090596	-	-
LBN-1.05	18/07/2015	Arran_LBN_2015_07_18.mts	00:20:42	00:22:29	13:26:28	13:28:15	55.546198	-5.090596	55.546068	-5.090829	-	-
LBN-1.06	18/07/2015	Arran_LBN_2015_07_18.mts	00:22:29	00:28:40	13:28:15	13:34:26	55.546068	-5.090829	55.545708	-5.092135	-	12.0
D3-1	19/07/2015	Arran_D3-1_2015_07_19.mts	00:01:14	00:32:02	08:55:47	09:26:35	55.616012	-5.127591	55.625517	-5.130281	6.3	9.3
D3-2.01	19/07/2015	Arran_D3-2_2015_07_19.mts	00:02:52	00:09:25	09:33:16	09:39:49	55.626215	-5.126380	55.624490	-5.125679	31.5	-
D3-2.02	19/07/2015	Arran_D3-2_2015_07_19.mts	00:09:25	00:20:44	09:39:49	09:51:08	55.624490	-5.125679	55.621371	-5.124277	-	-
D3-2.03	19/07/2015	Arran_D3-2_2015_07_19.mts	00:20:44	00:24:50	09:51:08	09:55:14	55.621371	-5.124277	55.620381	-5.123838	-	37.2
D3-3.01	19/07/2015	Arran_D3-3_2015_07_19.mts	00:01:16	00:03:40	10:02:08	10:04:32	55.619110	-5.129127	55.619322	-5.128259	8.3	-
D3-3.02	19/07/2015	Arran_D3-3_2015_07_19.mts	00:03:40	00:08:00	10:04:32	10:08:51	55.619322	-5.128259	55.619514	-5.126089	-	-
D3-3.03	19/07/2015	Arran_D3-3_2015_07_19.mts	00:08:00	00:10:14	10:08:51	10:11:06	55.619514	-5.126089	55.619647	-5.125079	-	-
D3-3.04	19/07/2015	Arran_D3-3_2015_07_19.mts	00:10:14	00:12:09	10:11:06	10:13:01	55.619647	-5.125079	55.619731	-5.124315	-	-
D3-3.05	19/07/2015	Arran_D3-3_2015_07_19.mts	00:12:09	00:19:39	10:13:01	10:20:31	55.619731	-5.124315	55.620112	-5.120872	-	-
D3-3.06	19/07/2015	Arran_D3-3_2015_07_19.mts	00:19:39	00:26:10	10:20:31	10:27:03	55.620112	-5.120872	55.620302	-5.118107	-	57.4
D3-4	19/07/2015	Arran_D3-4_2015_07_19.mts	00:01:57	00:39:20	10:41:40	11:19:03	55.616452	-5.126346	55.608542	-5.122117	23.0	21.6
D3-5.01	19/07/2015	Arran_D3-5_2015_07_19.mts	00:03:23	00:07:34	11:27:07	11:31:18	55.606568	-5.118008	55.607782	-5.118352	26.6	-
D3-5.02	19/07/2015	Arran_D3-5_2015_07_19.mts	00:07:34	00:12:18	11:31:18	11:36:02	55.607782	-5.118352	55.609145	-5.118683	-	-
D3-5.03	19/07/2015	Arran_D3-5_2015_07_19.mts	00:12:18	00:15:41	11:36:02	11:39:25	55.609145	-5.118683	55.610109	-5.119205	-	-
D3-5.04	19/07/2015	Arran_D3-5_2015_07_19.mts	00:15:41	00:17:19	11:39:25	11:41:03	55.610109	-5.119205	55.610543	-5.119397	-	-
D3-5.05	19/07/2015	Arran_D3-5_2015_07_19.mts	00:17:19	00:30:57	11:41:03	11:54:41	55.610543	-5.119397	55.614094	-5.121335	-	-
D3-5.06	19/07/2015	Arran_D3-5_2015_07_19.mts	00:30:57	00:31:37	11:54:41	11:55:21	55.614094	-5.121335	55.614214	-5.121425	-	-
D3-5.07	19/07/2015	Arran_D3-5_2015_07_19.mts	00:31:37	00:34:56	11:55:21	11:58:40	55.614214	-5.121425	55.615061	-5.121974	-	30.3
D3-6	19/07/2015	Arran_D3-6_2015_07_19.mts	00:03:20	00:12:32	12:12:13	12:21:25	55.613881	-5.117268	55.612163	-5.116302	46.2	47.6
D3-7.01	19/07/2015	Arran_D3-7_2015_07_19.mts	00:02:16	00:32:27	15:37:30	16:07:41	55.613918	-5.125551	55.605946	-5.125154	8.3	-
D3-7.02	19/07/2015	Arran_D3-7_2015_07_19.mts	00:32:27	00:42:07	16:07:41	16:17:21	55.605946	-5.125154	55.604167	-5.124464	-	19.0
T5-1	20/07/2015	Arran_T5-1_2015_07_20.mts	00:06:03	00:32:20	06:55:35	07:21:52	55.509186	-5.044909	55.510760	-5.042946	69.5	71.7
T5-2	20/07/2015	Arran_T5-2_2015_07_20.mts	00:05:21	00:49:01	07:40:54	08:24:34	55.515931	-5.031419	55.507876	-5.027751	73.8	83.9
T5-3.01	20/07/2015	Arran_T5-3_2015_07_20.mts	00:04:49	00:53:35	09:03:05	09:51:51	55.514850	-5.043516	55.509318	-5.042506	76.9	-
T5-3.02	20/07/2015	Arran_T5-3_2015_07_20.mts	00:53:35	01:10:07	09:51:51	10:08:23	55.509318	-5.042506	55.507633	-5.041133	-	72.5
T3-V01	17/09/2015	Arran_T3-V01_2015_09_17.m2ts	00:00:58	00:43:53	07:53:59	08:36:53	55.444640	-5.386407	55.450408	-5.388413	49.1	53.7
T3-V02	17/09/2015	Arran_T3-V02_2015_09_17.m2ts	00:00:08	00:51:58	08:58:04	09:49:54	55.444027	-5.380167	55.451715	-5.384803	46.0	54.2
T3-V03.01	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	00:00:11	00:34:42	11:09:19	11:43:50	55.446128	-5.370068	55.450692	-5.369050	47.6	45.3
T3-V03.02	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	00:34:42	00:35:26	11:43:50	11:44:34	55.450692	-5.369050	55.450848	-5.369102	45.3	46.8
T3-V03.03	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	00:35:26	00:53:26	11:44:34	12:02:34	55.450848	-5.369102	55.453857	-5.368757	46.8	44.7
T3-V03.04	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	00:53:26	00:53:58	12:02:34	12:03:06	55.453853	-5.368758	55.453903	-5.368720	44.7	44.6
T3-V03.05	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	00:53:58	01:02:15	12:03:06	12:11:23	55.453903	-5.368720	55.455315	-5.368480	44.6	42.0
T3-V03.06	17/09/2015	Arran_T3-V03_2015_09_17.m2ts	01:02:15	01:03:47	12:11:23	12:12:55	55.455318	-5.368480	55.455565	-5.368408	42.0	43.4
T3-V04	17/09/2015	Arran_T3-V04_2015_09_17.m2ts	00:00:00	00:52:04	12:33:39	13:25:43	55.445957	-5.375045	55.454650	-5.376225	52.6	53.1
D4-V01.01	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:00:09	00:05:54	14:03:55	14:09:38	55.420110	-5.306377	55.420167	-5.305627	29.9	29.2
D4-V01.02	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:05:54	00:11:54	14:09:38	14:15:38	55.420167	-5.305627	55.420060	-5.304132	29.2	29.9
D4-V01.03	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:11:54	00:15:26	14:15:38	14:19:10	55.420062	-5.304142	55.420128	-5.302973	29.9	30.0
D4-V01.04	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:15:26	00:23:13	14:19:10	14:26:57	55.420128	-5.302973	55.419892	-5.301085	30.0	28.7
D4-V01.05	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:23:13	00:35:29	14:26:57	14:39:13	55.419892	-5.301085	55.419572	-5.297808	28.7	29.4
D4-V01.06	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:35:29	00:41:07	14:39:13	14:44:51	55.419572	-5.297808	55.419490	-5.296178	29.4	30.2
D4-V01.07	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:41:07	00:46:36	14:44:51	14:50:20	55.419490	-5.296178	55.419442	-5.294457	30.2	29.9
D4-V01.08	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:46:36	00:49:01	14:50:20	14:52:45	55.419442	-5.294462	55.419393	-5.293757	29.9	31.4
D4-V01.09	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:49:01	00:50:17	14:52:45	14:54:01	55.419393	-5.293757	55.419360	-5.293368	31.4	31.5
D4-V01.10	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:50:17	00:51:32	14:54:01	14:55:16	55.419360	-5.293373	55.419352	-5.292962	31.5	31.7
D4-V01.11	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:51:32	00:52:32	14:55:16	14:56:16	55.419352	-5.292952	55.419358	-5.292623	31.7	32.1
D4-V01.12	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:52:32	00:53:36	14:56:16	14:57:20	55.419358	-5.292623	55.419352	-5.292247	32.1	32.6
D4-V01.13	17/09/2015	Arran_D4-V01_2015_09_17.m2ts	00:53:36	01:13:09	14:57:20	15:16:53	55.419352	-5.292247	55.419505	-5.284240	32.6	32.3
D4-V02.01	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:00:00	00:11:17	06:34:50	06:46:07	55.427285	-5.286310	55.427453	-5.289663	25.6	24.0
D4-V02.02	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:11:17	00:18:02	06:46:07	06:52:52	55.427453	-5.289663	55.427617	-5.291972	24.0	25.3
D4-V02.03	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:18:02	00:21:47	06:52:52	06:56:37	55.427617	-5.291975	55.427620	-5.293012	25.3	25.7
D4-V02.04	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:21:47	00:26:23	06:56:37	07:01:13	55.427618	-5.293008	55.427667	-5.294520	25.7	26.7
D4-V02.05	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:26:23	00:46:01	07:01:13	07:20:51	55.427667	-5.294523	55.427803	-5.300930	26.7	24.3
D4-V02.06	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:46:01	00:57:36	07:20:51	07:32:26	55.427803	-5.300930	55.427837	-5.304705	24.3	25.6
D4-V02.07	18/09/2015	Arran_D4-V02_2015_09_18.m2ts	00:57:36	01:03:29	07:32:26	07:38:19	55.427837	-5.304705	55.427832	-5.306565	25.6	26.8
D4-V03	18/09/2015	Arran_D4-V03_2015_09_18.m2ts	00:02:54	01:02:56	08:02:50	09:02:51	55.416615	-5.286163	55.416380	-5.307443	35.1	36.0
T2-V01	18/09/2015	Arran_T2-V01_2015_09_18.m2ts	00:00:05	00:36:57	09:50:38	10:27:30	55.398778	-5.188022	55.402613	-5.197888	53.2	55.7
T4-V01	18/09/2015	Arran_T4-V01_2015_09_18.m2ts	00:00:16	01:03:07	11:26:09	12:28:59	55.428792	-5.051575	55.436935	-5.030387	120.1	126.2
T4-V02	18/09/2015	Arran_T4-V02_2015_09_18.m2ts	00:01:02	00:30:24	12:59:37	13:28:59	55.428028	-5.045925	55.432317	-5.036638	126.9	125.2
T4-V03	18/09/2015	Arran_T4-V03_2015_09_18.m2ts	00:00:10	00:29:37	13:51:17	14:20:43	55.433727	-5.046907	55.438783	-5.036937	119.6	124.9
D6-V01.01	18/09/2015	Arran_D6-V01_2015_09_18.m2ts	00:00:13	00:23:27	15:00:09	15:23:19	55.435267	-5.085708	55.431268	-5.092735	42.7	37.5
D6-V01.02	18/09/2015	Arran_D6-V01_2015_09_18.m2ts	00:23:27	00:26:09	15:23:19	15:26:02	55.431268	-5.092735	55.430852	-5.093747	37.5	34.1
D6-V02	19/09/2015	Arran_D6-V02_2015_09_19.m2ts	00:00:13	01:01:25	06:48:36	07:49:46	55.428192	-5.108553	55.440272	-5.088433	27.7	36.4
D6-V03.01	19/09/2015	Arran_D6-V03_2015_09_19.m2ts	00:00:18	00:06:36	08:09:19	08:15:37	55.429488	-5.109865	55.431293	-5.109477	21.9	17.2
D6-V03.02	19/09/2015	Arran_D6-V03_2015_09_19.m2ts	00:06:36	00:22:20	08:15:37	08:31:21	55.431293	-5.109477	55.434900	-5.105692	17.2	14.2
D6-V03.03	19/09/2015	Arran_D6-V03_2015_09_19.m2ts	00:22:20	00:42:13	08:31:21	08:51:14	55.434900	-5.105692	55.439522	-5.099095	14.2	15.3
D6-V03.04	19/09/2015	Arran_D6-V03_2015_09_19.m2ts	00:42:13	00:48:24	08:51:14	08:57:25	55.439522	-5.099095	55.440837	-5.096958	15.3	16.5
D6-V03.05	19/09/2015	Arran_D6-V03_2015_09_19.m2ts	00:48:24	00:58:38	08:57:25	09:07:39	55.440837	-5.096958	55.443080	-5.093222	16.5	17.0
D6-V04.01	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:00:13	00:22:39	09:28:49	09:51:15	55.425672	-5.106730	55.428407	-5.098892	32.0	28.9
D6-V04.02	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:22:39	00:22:58	09:51:15	09:51:34	55.428407	-5.098892	55.428425	-5.098765	28.9	29.1
D6-V04.03	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:22:58	00:33:51	09:51:34	10:02:27	55.428425	-5.098765	55.430525	-5.094775	29.1	29.2
D6-V04.04	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:33:51	00:34:50	10:02:27	10:03:26	55.430525	-5.094775	55.430782	-5.094450	29.2	28.7
D6-V04.05	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:34:50	00:35:23	10:03:26	10:03:59	55.430775	-5.094453	55.430923	-5.094327	28.7	28.2
D6-V04.06	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:35:23	00:36:54	10:03:59	10:05:30	55.430923	-5.094327	55.431348	-5.093920	28.2	27.5
D6-V04.07	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:36:54	00:39:30	10:05:30	10:08:06	55.431348	-5.093920	55.432090	-5.093367	27.5	27.1
D6-V04.08	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:39:30	00:40:30	10:08:06	10:09:06	55.432090	-5.093367	55.432360	-5.093088	27.1	27.4
D6-V04.09	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:40:30	00:41:32	10:09:06	10:10:08	55.432357	-5.093092	55.432600	-5.092887	27.4	28.7
D6-V04.10	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:41:32	00:43:24	10:10:08	10:12:00	55.432605	-5.092882	55.433035	-5.092522	28.7	35.1
D6-V04.11	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:43:24	00:44:11	10:12:00	10:12:47	55.433035	-5.092522	55.433230	-5.092383	35.1	37.0
D6-V04.12	19/09/2015	Arran_D6-V04_2015_09_19.m2ts	00:44:11	00:57:29	10:12:47	10:26:05	55.433225	-5.092388	55.436345	-5.089642	37.0	38.3
T4-V04	19/09/2015	Arran_T4-V04_2015_09_19.m2ts	00:00:11	00:30:28	11:20:36	11:50:53	55.442315	-5.031408	55.437920	-5.036852	125.0	124.8
T4-V05	19/09/2015	Arran_T4-V05_2015_09_19.m2ts	00:01:25	01:11:08	12:17:13	13:26:56	55.424685	-5.041435	55.438487	-5.025355	114.2	117.1
T2-V02	19/09/2015	Arran_T2-V02_2015_09_19.m2ts	00:00:14	00:41:09	14:39:47	15:20:42	55.389178	-5.200332	55.394902	-5.213377	55.7	55.7
T2-V03	19/09/2015	Arran_T2-V03_2015_09_19.m2ts	00:00:10	00:34:54	16:05:11	16:39:55	55.394257	-5.194598	55.399192	-5.207360	55.9	56.1
T2-V04	21/09/2015	Arran_T2-V04_2015_09_21.m2ts	00:00:09	00:39:13	07:30:55	08:09:59	55.410658	-5.170963	55.416127	-5.185095	46.7	44.2
T2-V05	21/09/2015	Arran_T2-V05_2015_09_21.m2ts	00:00:12	00:37:27	08:28:46	09:06:01	55.407558	-5.177010	55.412803	-5.189225	52.8	44.8
T2-V06	21/09/2015	Arran_T2-V05_2015_09_21.m2ts	00:00:09	00:42:45	09:22:56	10:05:32	55.403028	-5.180425	55.408453	-5.194780	53.8	48.1
D6-V05.01	21/09/2015	Arran_D6-V05_2015_09_21.m2ts	00:00:01	00:26:26	11:14:34	11:40:59	55.422880	-5.103743	55.427460	-5.096212	38.9	42.3
D6-V05.02	21/09/2015	Arran_D6-V05_2015_09_21.m2ts	00:26:26	00:55:20	11:40:59	12:09:53	55.427460	-5.096212	55.432980	-5.087252	42.3	39.9
D6-V05.03	21/09/2015	Arran_D6-V05_2015_09_21.m2ts	00:55:20	00:57:42	12:09:53	12:12:15	55.432980	-5.087252	55.433493	-5.086475	39.9	40.9
D6-V05.04	21/09/2015	Arran_D6-V05_2015_09_21.m2ts	00:57:42	01:03:20	12:12:15	12:17:53	55.433490	-5.086477	55.434980	-5.084582	40.9	43.7
D4-V04.01	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:00:06	00:00:31	13:35:27	13:35:52	55.423938	-5.306852	55.423955	-5.306713	29.7	29.7
D4-V04.02	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:00:31	00:12:44	13:35:52	13:48:05	55.423955	-5.306713	55.424037	-5.302240	29.7	28.8
D4-V04.03	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:12:44	00:19:26	13:48:05	13:54:47	55.424037	-5.302230	55.423458	-5.299667	28.8	29.1
D4-V04.04	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:19:26	00:24:59	13:54:47	14:00:20	55.423458	-5.299667	55.422827	-5.297450	29.1	29.0
D4-V04.05	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:24:59	00:29:53	14:00:20	14:05:14	55.422827	-5.297450	55.422467	-5.295505	29.0	28.3
D4-V04.06	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:29:53	00:30:29	14:05:14	14:05:50	55.422467	-5.295505	55.422473	-5.295263	28.3	28.5
D4-V04.07	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:30:29	00:34:52	14:05:50	14:10:13	55.422473	-5.295263	55.422617	-5.293703	28.5	27.9
D4-V04.08	21/09/2015	Arran_D4-V04_2015_09_21.m2ts	00:34:52	00:49:29	14:10:13	14:24:50	55.422617	-5.293703	55.422910	-5.288487	27.9	29.4
D4-V05.01	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:00:19	00:20:09	14:45:22	15:05:12	55.413662	-5.303665	55.413958	-5.295830	36.4	32.5
D4-V05.02	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:20:09	00:20:44	15:05:12	15:05:47	55.413958	-5.295830	55.413947	-5.295613	32.5	32.6
D4-V05.03	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:20:44	00:23:14	15:05:47	15:08:17	55.413943	-5.295603	55.413958	-5.294692	32.6	32.7
D4-V05.04	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:23:14	00:26:10	15:08:17	15:11:13	55.413958	-5.294692	55.414038	-5.293545	32.7	32.9
D4-V05.05	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:26:10	00:31:02	15:11:13	15:16:05	55.414038	-5.293545	55.414065	-5.291850	32.9	33.7
D4-V05.06	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:31:02	00:31:28	15:16:05	15:16:31	55.414065	-5.291850	55.414063	-5.291690	33.7	33.6
D4-V05.07	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:31:28	00:32:08	15:16:31	15:17:11	55.414063	-5.291690	55.414055	-5.291468	33.6	33.8
D4-V05.08	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:32:08	00:32:30	15:17:11	15:17:33	55.414055	-5.291468	55.414055	-5.291358	33.8	33.7
D4-V05.09	21/09/2015	Arran_D4-V05_2015_09_21.m2ts	00:32:30	00:39:02	15:17:33	15:24:05	55.414055	-5.291358	55.414085	-5.289220	33.7	34.7

Annex 2: Physical and biological descriptions of the survey sites, with indicators of conservation importance of habitats and species

Sample codes correspond with those in Annex 1. Priority Marine Feature (PMF) codes (unitalicized font) and Protected Feature (PF) codes (bold font) and Habitat Directive Annex I codes are defined in Annex 4. Uncertain biotope assignments are italicized.

Sample	Substrate	Biota	Biotope	Annex I habitat	PF/PMF
T1-1	Mud (100%) with sparsely scattered cobbles (<1%), boulders (<1%) and possibly small bedrock outcrop (<1%).	Mud moderately densely burrowed, particularly by Calocaris macandreae (C) and Nephrops norvegicus (C, 79 animals seen). Rock with hydroids (R overall). Pennatula phosphorea (R), Urticina sp.? (R), Caridea sp. (O), Munida spp. (R), Paguridae spp. (R) including Pagurus bernhardus, Asterias rubens (F), small teleost spp. (R), Glyptocephalus cyanoglossus? (P), Lumpenus lampretaeformis (P). Dense euphausiids in water column at start of run.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-2	Mud (100%).	Very poor visibility and no photos. Mud moderately densely burrowed, particularly by Calocaris macandreae (C) and Nephrops norvegicus (probably C, 10 animals seen). Caridea sp. (O), teleost spp. (R), small Glyptocephalus cyanoglossus? (P). Dense euphausiids in water column at start of run.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-3.01	Sandy mud (99%) and shell gravel (1%) with sparse cobbles (<1%) and boulders (<1%).	Moderate density of megafaunal burrows including Calocaris macandreae (F, locally C) and Nephrops norvegicus (C, 13 animals seen). Pennatula sp.? (R), Sabella pavonina tubes (R), Munida spp. (O), Paguridae spp. (O), Caridea sp. (P), Turritella communis shells (R, occupancy unknown), Asterias rubens (F), teleost spp. (O), Glyptocephalus cyanoglossus? (O).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-3.02	Mud or sandy mud (87%) with shell gravel (1%) and scattered pebbles (2%), cobbles (8%) and boulders (2%).	Silted stones with short hydroid turf (C). Sediment with very sparse, small burrows (R). Munida spp. (O), Paguridae sp. (P), Asterias rubens (P).	SS.SMu.CFiMu, CR.LCR	RF:ST	-
T1-3.03	Mud (100%) with large boulder (<1%).	Moderate density of megafaunal burrows including Calocaris macandreae (F, locally C) and Nephrops norvegicus (C, 17 animals seen). Virgularia mirabilis (R), Cerianthus lloydii (R), Aphrodita aculeata (P), Sabella pavonina tubes (R), Paguridae spp. (O), Caridea sp. (O), Asterias rubens (O), teleost spp. (O), Glyptocephalus cyanoglossus? (P), Lumpenus lampretaeformis (P), Gadidae spp. (O). Boulder with hydroids (where C), Parasmittina trispinosa? (R) and small Actiniaria sp.? (where C). Euphausiids in water column.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-4	Sandy mud (99%) and shell gravel (1%) with sparse cobbles (<1%) and boulders (<1%).	Moderate density of megafaunal burrows including Calocaris macandreae (F, locally C) and Nephrops norvegicus (C, 29 animals seen). Hydroids (R, locally A on stones), Sabella pavonina tubes (R), emergent infaunal tubes (P), Munida sp. (R), Paguridae spp. (O), Caridea sp. (P), Buccinum undatum? (P), Turritella communis shells (R, occupancy unknown), Asterias rubens (F), teleost spp. (O), Glyptocephalus cyanoglossus? (O), Lumpenus lampretaeformis (O). Dense euphausiids in water column.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-5	Mud (100%).	Moderate density of megafaunal burrows including Calocaris macandreae (C, at least locally) and Nephrops norvegicus (C, 32 animals seen). Sabella pavonina tubes? (R), Caridea sp. (P), Asterias rubens (O), teleost spp. (O), Glyptocephalus cyanoglossus? (O), Lumpenus lampretaeformis (P). Faint parallel scars, possibly trawl disturbance (00:13:52 - 00:13:55). Euphausiids in water column.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D5-1.01	Sandy mud or possibly mud (99%) with shell gravel and broken shell material (1%).	Shell material with hydroids (R) and Omalosecosa ramulosa (R). Sediment with worm casts and emergent infaunal tubes and lightly burrowed by megafauna, except for Nephrops norvegicus (C, 5 animals seen). Munida rugosa (F), Aequipecten opercularis (R), Echinus esculentus (P), teleost spp. (O), Pleuronectiformes sp. (P). Gradual transition to following biotope.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D5-1.02	Sandy mud (99%) with shell gravel and broken shell material (1%).	Sediment with worm casts, small holes, emergent infaunal tubes and very sparse megafaunal burrows. Nemertesia ramosa (P), Munida rugosa (F), Paguridae spp. in Turritella communis shells (O), teleost sp. (P), Porania pulvillus (P), Pleuronectiformes sp. (P). Gradual transitions to preceding and following biotopes.	SS.SMu.CSaMu	-	-
D5-1.03	Muddy sand (88%) with shell gravel (5%), stone gravel (5%), shells (1%), pebbles (1%) and cobbles (<1%).	Sediment with Cerianthus lloydii (F), small holes, emergent infaunal tubes and small burrow in transitional zone. Shell and stone material with hydroids (O), Buccinum undatum egg mass, Omalosecosa ramulosa (R) and Corella parallelogramma (O). Munida rugosa (F), Aequipecten opercularis (O), Porania pulvillus (O), Callionymus sp. (P), Gobiidae sp. (P), small teleost sp. (P).	SS.SSa.CMuSa	-	-
D5-1.04	Mixed substrate of muddy sand (66%) with shell gravel (15%) and stone gravel (5%), shells and large shell material (2%), pebbles (5%), cobbles (5%) and boulders (1%).	Stones encrusted with pink coralline algae (R), serpulid worms (P), Balanus spp. (P) and Parasmittina trispinosa? (R) and supporting hydroids (F) including Nemertesia ramosa (O) and N antennina (F), Omalosecosa ramulosa (F), Bugula sp. (P) and solitary ascidians (O) including Corella parallelogramma (O) and Ascidia virginea (O). Sediment with small holes, emergent infaunal tubes and Cerianthus lloydii (A). Munida rugosa (O), Liocarcinus sp. (P), Aequipecten opercularis (O), Pecten maximus (P), Asterias rubens (F), Astropecten irregularis (P), Porania pulvillus (F), small patch of Ophiocomina nigra (where A), Echinus esculentus (P).	SS.SMx.CMx.ClloMx.Nem	-	-
D5-2.01	Sandy mud or cohesive muddy sand (98%) with gravel (2%), shells (<1%), pebbles (<1%), cobbles (<1%) and boulders (<1%).	Sediment with worm casts, small holes, emergent infaunal tubes and very sparse megafaunal burrows. Hydroids (O, locally A on boulders) including Nemertesia ramosa (P), Aphrodita aculeata? (P), Munida rugosa (F), Paguridae spp. (O), Liocarcinus depurator (P), Aporrhais pespelicani shells (P, but shell occupancy uncertain), Buccinum undatum? (P), Aequipecten opercularis (O), teleost sp. (P), Porania pulvillus (O), Corella parallelogramma (P), single clump of Ascidiella aspersa. Gradual transitions to following biotope.	SS.SMu.CSaMu	-	-
D5-2.02	Muddy sand (98%) with gravel (1%), pebbles (1%), shells (<1%) and cobbles (<1%).	Sediment with Cerianthus lloydii? (O), small holes and emergent infaunal tubes. Shell and stone material with hydroids (O) including Nemertesia ramosa (O), Corella parallelogramma (P) and Ascidiella aspersa (O). Munida rugosa (F), Liocarcinus spp. (O) including L. depurator, Aporrhais pespelicani shell (occupancy unknown), Aequipecten opercularis (F). Boundaries with previous and following biotopes uncertain.	SS.SSa.CMuSa	-	-
D5-2.03	Mixed substrate of muddy sand (53%) with shell gravel (20%) and stone gravel (10%), shells and large shell material (2%), pebbles (10%) and cobbles (5%).	Stones encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydroids (F) including Nemertesia ramosa (F) and N antennina (F), Omalosecosa ramulosa (F), Alcyonidium diaphanum (P) and solitary ascidians (F) including Corella parallelogramma (F) and Ascidia virginea (O). Sediment with emergent infaunal tubes including Chaetopterus variopedatus? (P), and Cerianthus lloydii (A). Munida rugosa (O), Paguridae sp. (P), Cancer pagurus (P), Buccinum undatum egg mass, mollusc egg ribbon, Aequipecten opercularis (O), Pecten maximus (P), Marthasterias glacialis (F), Crossaster papposus (P), Asterias rubens (F), Astropecten irregularis (P), Luidia ciliaris (P), Porania pulvillus (F), Echinus esculentus (F).	SS.SMx.CMx.ClloMx.Nem	-	-
D5-2.04	Highly mixed substrate of silty sand (20%) and gravel (35%) and pebbles (30%), with cobbles (10%) and boulders (5%).	Stones encrusted with pink coralline algae (R), serpulid worms, Balanus spp. (P) and Parasmittina trispinosa? (R) and supporting hydroids (F, A-S on boulders) including Nemertesia ramosa (F) and N. antennina (P), mollusc egg strings, Omalosecosa ramulosa (F), Alcyonidium diaphanum (P), as well as solitary ascidians (C, locally A) including Ascidia virginea (F), A. mentula, Ascidiella aspersa (locally A), Corella parallelogramma (F) and Ciona intestinalis (locally A). Cerianthus lloydii (C initially in transitional area), Munida rugosa (O), Paguridae spp. (O), Antedon spp. (F), Crossaster papposus (P), Asterias rubens (C), Porania pulvillus (F), Marthasterias glacialis (F), patches of Ophiocomina nigra (locally A), Echinus esculentus (F), small teleost spp. (O). Considered a mosaic of AntAsH on boulders and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs.AntAsH	RF:ST	-
D5-2.05	Mixed substrate of silty sand (60%) with gravel (25%), pebbles (10%) and cobbles (5%).	Stones encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydroids (O), Omalosecosa ramulosa (O), and solitary ascidians (O) including Ascidia virginea (P). Sediment with Chaetopterus variopedatus? (P), and Cerianthus lloydii (O). Munida rugosa (P), Asterias rubens (C), Porania pulvillus (O), Callionymus lyra (P).	SS.SMx.CMx	-	-
D5-2.06	Muddy sand (92%) with shell and stone gravel (5%), pebbles (2%) and shells (1%).	Sediment with small holes and mounds. Stones and shells with hydroids (O) including Nemertesia ramosa and encrusted with serpulid worms (R). Sparse Turritella communis shells (occupancy unknown), Porania pulvillus (O).	SS.SSa.CMuSa	-	-
D5-2.07	Cohesive muddy sand (97%) with shell and stone gravel (2%), and shells (1%); boulder (<1%).	Sediment with small holes, mounds, worm casts, emergent infaunal tubes including Chaetopterus variopedatus?, and very sparse megafaunal burrows. Hydroids (O, locally A on boulder) including Nemertesia ramosa and N. antennina, Aphrodita aculeata (P), Munida rugosa (F), small Brachyura sp. (P), Liocarcinus sp. (P), Turritella communis shells (occupancy generally unknown, although some occupied by Paguridae spp.), Aporrhais pespelicani shells (O, occupancy unknown), Buccinum undatum (P), Asterias rubens (F), Porania pulvillus (P), Corella parallelogramma (P), Callionymus sp. (P).	SS.SMu.CSaMu	-	-
D5-2.08	Cohesive muddy sand or sandy mud (99%) with shell gravel and broken shell material (1%); shells (<1%).	Shell material with hydroids (O) including Nemertesia ramosa (O) and N. antennina (O). Sediment with small holes, small mounds, worm casts, Terebellidae sp. tentacles (P) and emergent infaunal tubes including Chaetopterus variopedatus? and lightly burrowed by megafauna, except for Nephrops norvegicus (C, 1 animals seen). Munida rugosa (F), Liocarcinus sp. (P), Turritella communis shells (P, occupancy unknown), Aporrhais pespelicani shells (occupancy unknown), Asterias rubens (P), Pleuronectiformes sp. (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D5-2.09	Cohesive muddy sand or sandy mud (85%) with boulders (5%) and bedrock cliff (10%).	Sediment with emergent infaunal tubes, small mounds and lightly burrowed by megafauna. Silted stones and bedrock with patchy hydroid turf (locally S) including Nemertesia ramosa F), Alcyonium digitatum? (R), sparse serpulid worms (P) and solitary ascidians (O). Paguridae spp. (O), Munida rugosa (F), Turritella communis shells (occupancy unknown).	SS.SMu.CFiMu.SpnMeg, CR.LCR.BrAs	RF:BR	BM:SB
D5-2.10	Cohesive muddy sand or sandy mud (98%) with shell gravel and broken shell material (2%).	Sediment with small holes and emergent infaunal tubes including Chaetopterus variopedatus? and lightly burrowed by megafauna, except for Nephrops norvegicus (C, possibly 1 animals seen). Hydroids (O), Munida rugosa (F), Turritella communis shells (P, occupancy unknown but probably all empty), Porania pulvillus (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D5-3.01	Mixed substrate of muddy sand (25%) with maerl gravel (20%), shell and stone gravel (25%), shells (10%), pebbles (15%), cobbles (5%) and boulders (<1%).	Stones and shells encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydroids (F, locally S on boulders) including Nemertesia ramosa (F) and N antennina (O), solitary ascidians (F, locally A) including Ascidiella aspersa (F, locally A in clumps) and Ascidia mentula (P), and filiform red algae (R). Drift algae also present, although some Desmarestia spp (R) and Saccharina latissima (O) may be living; latter with Membranipora membranacea and Electra pilosa. Sediment with emergent infaunal tubes and Cerianthus lloydii (C, locally A) and with very sparse megafaunal burrows, one containing Nephrops norvegicus. Munida rugosa (F), Liocarcinus spp. (P), mollusc egg ribbon, Pecten maximus? (P), Marthasterias glacialis (P), Crossaster papposus (F), Asterias rubens (F), Astropecten irregularis (P), Porania pulvillus (O), Henricia sp. (P), Echinus esculentus (O).	SS.SMx.CMx.ClloMx.Nem	SB:MX	-
D5-3.02	Mixed substrate of muddy sand (63%), with gravel (25%), pebbles (10%), cobbles (<1%) and shells (2%).	Stones and shells with hydroids (O), serpulid worms (P), Ascidiella aspersa (C, often in clumps) and filiform red algae (R). Sediment with small holes and sparse Cerianthus lloydii (P). Drift algae, Munida rugosa (F), Liocarcinus spp. (F), Aequipecten opercularis (O), Porania pulvillus (P), small teleost sp. (P).	SS.SMx.CMx	-	-
D5-3.03	Muddy sand (88%) with gravel (10%), shells (1%) and pebbles (1%).	Hydroids (O), Munida rugosa (F), Pagurus bernhardus (P), Liocarcinus spp. (O), Aequipecten opercularis (P), Porania pulvillus (P), Ascidiella aspersa (C), Callionymus sp. (P), drift kelp. Boundaries with previous and following biotopes gradual and ill-defined.	SS.SSa.CMuSa	-	-
D5-3.04	Cohesive muddy sand (93%) with gravel (5%) and shells (2%); boulder (<1%).	Sediment with small holes, small mounds, worm casts and emergent infaunal tubes. Hydroids (O) including Nemertesia antennina? (P), Munida rugosa (F), Turritella communis shells (occupancy generally unknown, although at least one occupied by Paguridae sp.), Aporrhais pespelicani shells (O, occupancy unknown), Aequipecten opercularis (O), Marthasterias glacialis? (P), Porania pulvillus (O), Ascidiella aspersa (F), Callionymus spp. (O), small teleost sp. (P), drift algae, though some filiform red algae may be functional (R). Boundary with previous biotope gradual and ill-defined.	SS.SMu.CSaMu	-	-
D5-4.01	Maerl (65%) and stone (20%) gravel with shells (15%).	Saccharina latissima (C) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa. Patchy turf of filiform red algae (C) including Bonnemaisonia asparagoides?, with brown algae including Desmarestia ligulata, D. aculeata and Chorda filum (P). Shells encrusted with serpulid worms (P). Sediment with Cerianthus lloydii (C). Marthasterias glacialis (C), Ascidiella aspersa (C, at least locally).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D5-4.02	Mixed substrate of muddy sand (40%) with maerl gravel (15%), shell and stone gravel (20%), shells (10%), pebbles (15%), cobbles (<1%) and boulders (<1%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P) and supporting hydroids (O, locally A on boulders) including Nemertesia ramosa (O) and N antennina? (P), solitary ascidians (A) including Ascidiella aspersa (A) and Corella parallelogramma (P). Drift algae present but some algae apparently functional including filiform red algae (R), foliose red algae (R) and Saccharina latissima (O) with Membranipora membranacea and Electra pilosa. Sediment with small holes, emergent infaunal tubes and Cerianthus lloydii (C). Munida rugosa (F), Paguridae sp. (P), Liocarcinus depurator (P), Inachus sp. (P), Cancer pagurus (P), Marthasterias glacialis (F), Crossaster papposus (P), Asterias rubens (P), Porania pulvillus (O), Henricia sp. (P).	SS.SMx.CMx.ClloMx.Nem	SB:MX	-
D5-4.03	Slightly mixed substrate but largely muddy sand (79%) with scattered gravel (15%, denser locally), pebbles (5%) and shells (1%); cobbles (<1%).	Sediment with small holes, worm casts, emergent infaunal tubes, Sabellidae sp. (P) and Cerianthus lloydii (F). Stones and shells support hydroids (F) including Nemertesia ramosa (F) and N. antennina (F), Corella parallelogramma (P), clumps of Ascidiella aspersa (C) and filiform red algae (R). Munida rugosa (F), Liocarcinus spp. (O) including L. depurator, Turritella communis shells (occupancy unknown), Asterias rubens (P), Henricia sp. (P), Porania pulvillus (P),	SS.SMx.CMx.ClloMx.Nem	SB:MX	-
D5-4.04	Muddy sand (93%) with gravel (5%) and shells and large shell material (2%); cobbles (<1%), boulders (<1%).	Sediment with small holes, polychaete casts, emergent infaunal tubes including possibly Chaetopterus variopedatus, Cerianthus lloydii (F), juvenile Virgularia mirabilis (O) and single megafaunal burrow implying cohesive sediment. Hydroids (F) including Nemertesia antennina (F) and N. ramosa (P), Munida rugosa (F), Paguridae sp. (P), Liocarcinus spp. (O) including L. depurator, Turritella communis shells (occupancy unknown), Porania pulvillus (O), Astropecten irregularis? (P), Luidia ciliaris (F), Ascidiella aspersa (F), Ciona intestinalis (P), small teleost sp. (P), filiform red algae (R), drift algae.	SS.SMu.CSaMu	-	-
D5-5.01	Maerl gravel (55%), stone and shell gravel (20%), shells (10%), pebbles (10%), muddy sand (5%).	Saccharina latissima (A, locally C) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa; Chorda filum (C). Patchy red algal turf (C) of filiform (C) and foliose (R) species including Bonnemaisonia asparagoides, brown algae including Desmarestia spp. (O-F) with D. aculeata (P) and D. viridis? (P), and Ulva sp. (R). Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P). Cerianthus lloydii (C, at least locally), Chaetopterus variopedatus (P), Marthasterias glacialis (C), Asterias rubens (F), Porania pulvillus (P), Astropecten irregularis? (P), Luidia ciliaris (P), Echinus esculentus (P), Ascidiella aspersa (A locally), small teleost spp. (P), live maerl thalli (R, <1%).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D5-5.02	Mixed substrate of muddy sand (20%) with maerl gravel (25%), shell and stone gravel (40%), shells (5%), pebbles (10%) and cobbles (<1%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P) and supporting hydroids (O) including Nemertesia ramosa (O) and N antennina (O), and Ascidiella aspersa (C, locally A). Filiform red algae (R), Saccharina latissima (F, locally A) with Membranipora membranacea and Electra pilosa, but probably drift material from adjacent biotope. Sediment with emergent infaunal tubes including Chaetopterus variopedatus (P), and Cerianthus lloydii (A). Munida rugosa (P), Inachus sp. (P), Marthasterias glacialis (F), Crossaster papposus (P), Asterias rubens (P), Luidia ciliaris (F), Porania pulvillus (O), small teleost sp. (P). Gradual transition with following biotope.	SS.SMx.CMx.ClloMx.Nem	SB:MX	-
D5-5.03	Muddy sand (94%) with scattered shell gravel (5%), shells (1%) and pebbles (<1%).	Sediment with small holes and emergent infaunal tubes and Cerianthus lloydii (F). Hydroids (F) including Nemertesia antennina (F), Munida rugosa (F), sparse Turritella communis shells (occupancy unknown), Ascidiella aspersa (F), Callionymus lyra (P), filiform red algae (R), drift kelp.	SS.SMu.CSaMu	-	-
D5-6.01	Maerl gravel (50%), stone and shell gravel (30%), shells (10%), pebbles (5%), muddy sand (5%).	Blanket of Saccharina latissima (A) with fronds supporting some dense mats of Membranipora membranacea and Electra pilosa; Chorda filum (C). Patchy red algal turf (C) of filiform species including Bonnemaisonia asparagoides, brown algae including Desmarestia spp. (O-F) with D. aculeata (P), Dictyota dichotoma (P), and Ulva sp. (R). Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P). Hydroids (P) including Nemertesia antennina, Cerianthus lloydii (C, at least locally), Chaetopterus variopedatus (P), bivalve siphons (P), Marthasterias glacialis (C), Asterias rubens (C), Luidia ciliaris (O), Neopentadactyla mixta (P), small teleost sp. (P), Gobiidae sp. (P), live maerl thalli (R, <1%). The density of some taxa may be underestimated due to the widespread blanket of kelp.	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D5-6.02	Mixed substrate of silty sand (30%) with maerl gravel (20%), shell and stone gravel (20%), shells (15%), pebbles (15%) and cobbles (<1%).	Stones and shells encrusted with pink coralline algae (R), serpulid worms (P) and Parasmittina trispinosa? (R), and supporting sparse hydroids (P) including Nemertesia ramosa and N. antennina, Ascidiella aspersa (F) and Ascidia mentula (O). Filiform red algae (R), Saccharina latissima (O) but probably drift material from adjacent biotope. Sediment with Chaetopterus variopedatus (P), and Cerianthus lloydii (A). Liocarcinus sp. (P), Pecten maximus (P), Asterias rubens (P), Astropecten irregularis? (O), Henricia sp. (P), Porania pulvillus (O), live maerl thalli (R, <1%).	SS.SMx.CMx.ClloMx	SB:MX	-
D1-1.01	Slightly silty maerl gravel (80%) with shells (10%), sand (5%) and stone gravel (5%).	Patchy algal turf (C) including filiform reds (F, locally C, possibly mostly Bonnemaisonia asparagoides), Desmarestia aculeata? (P), Dictyota dichotoma (P), Saccharina latissima (F) and Chorda filum (P). Cerianthus lloydii (A), serpulid worms (P), Cancer pagurus (P), Marthasterias glacialis (F), Asterias rubens (F), Astropecten irregularis (P), Porania pulvillus (R), pink encrusting coralline algae (R), live maerl (<1%, R).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-1.02	Changes from silty maerl gravel and sand with shells to silty gravelly sand with pebbles and shells to silty sandy gravel with shells, pebbles and cobbles.	Sediment with worm casts and Cerianthus lloydii (C, but A over large areas). Stones and shells with Metridium dianthus (P), serpulid worms (P), pink encrusting coralline algae, hydroids (O) including Halecium halecinum (P), Alcyonidium diaphanum? (R), solitary ascidians (R) and sparse erect algae including filiform reds (R), Desmarestia aculeata? (R) and Saccharina latissima (O). Munida rugosa (O), Atelecyclus rotundatus (R), Liocarcinus depurator (O), Cancer pagurus (F), Buccinum undatum (R), Pecten maximus (R), Aequipecten opercularis (O), Asterias rubens (O), Porania pulvillus (O), Luidia ciliaris (P), Scyliorhinus sp. (P), Callionymus spp. (R).	SS.SMx.CMx.ClloMx	SB:MX	-
D1-1.03	Mixed substrate of silty sand (35%) and gravel (35%) with pebbles (5%) and scattered clumps of silted boulders (20%) and cobbles (5%).	Larger stones encrusted with pink coralline algae (R) and supporting hydroid turf (where A), erect bryozoans (P) including Alcyonidium diaphanum, and solitary ascidians (F) including Ascidia virginea and Corella parallelogramma. Munida rugosa (P), Buccinum undatum eggs, Porania pulvillus (P), Luidia ciliaris (P).	SS.SMx.CMx, CR.LCR.BrAs	SB:MX, RF:ST	-
D1-1.04	Mixed substrate of silty sand (35%) and gravel (35%) with pebbles (5%) and scattered, silted boulders (20%) and cobbles (5%).	Poor visibility. Bed of Ophiocomina nigra (A, locally S). Boulders with pink encrusting coralline algae (R) and solitary ascidians (P) including Ascidia virginea. Crossaster papposus (P), Luidia ciliaris (P)	SS.SMx.CMx.OphMx	SB:MX	-
D1-2.01	Mixed coarse substrate of maerl gravel (30%) and stone gravel (35%) with pebbles (20%), shells (10%) and scattered boulders (<1%) and cobbles (5%).	Mostly dense cover of Saccharina latissima (A, locally C) with fronds supporting often dense mats of Membranipora membranacea and Electra pilosa. Understorey of foliose (P) and filiform (C) red algae including Bonnemaisonia asparagoides? and Phycodrys rubens?, Dictyota dichotoma and Desmarestia aculeata. Stones encrusted with pink coralline algae (O) and serpulid worms (P). Crossaster papposus (P), Asterias rubens (P), Marthasterias glacialis (F), Porania pulvillus (R), Echinus esculentus (F).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-2.02	Mixed coarse substrate of maerl gravel (40%), stone and shell gravel (30%), coarse sand (10%), pebbles (15%), shells (5%) and scattered boulders (<1%) and cobbles (<1%). Sediment becomes finer along run.	Dense cover of Saccharina latissima for part of run (F overall but A locally) with fronds supporting often dense mats of Membranipora membranacea and Electra pilosa. Understorey of foliose (P) and filiform (C) red algae including Bonnemaisonia asparagoides?, and Desmarestia aculeata. Stones encrusted with pink coralline algae (O), serpulid worms (P) and Balanus spp. and supporting hydroids. Sediment with Cerianthus lloydii (C locally) and bivalve siphons. Pecten maximus (R), nudibranch egg string, Asterias rubens (P), Marthasterias glacialis (F), Henricia sp. (P), Porania pulvillus (R), Luidia ciliaris (P).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-2.03	Mixed, silty, coarse substrate of maerl gravel (50%), stone gravel (20%), sand (15%), pebbles (7%), shells (8%) and scattered cobbles (<1%). Sediment becomes finer and less mixed along run.	Stones and shells encrusted with serpulid worms and pink coralline algae (R) and supporting digitiform orange sponge (R), hydroids (O) including Nemertesia antennina, Alcyonium digitatum (R), bryozoans (R), Corella parallelogramma (P) and filiform (R) and foliose (R) red algae; Saccharina latissima (O, probably drift material. Sediment with Cerianthus lloydii (A), Chaetopterus variopedatus? (P) and Neopentadactyla mixta (P, 1 individual seen). Inachus spp. (P), Aequipecten opercularis (R), Astropecten irregularis? (R), Porania pulvillus (R), Henricia spp. (P). Biotope gradually transitional at start and end.	SS.SCS.CCS.Nmix	SB:GS	MC
D1-2.04	A slightly mixed silty sand with gravel (15%), sand (80%), shells (5%) and cobbles (<1%).	Stones and shells with clumps of hydroids (O) including Nemertesia antennina, Balanus spp. (R), bryozoans (R), solitary ascidians (O) including Corella parallelogramma and filiform red algae (O, some possibly drift). Munida rugosa (O), Aequipecten opercularis (O), Asterias rubens (F), Henricia sp. (R). Sediment with small holes, worm casts and Cerianthus lloydii (initially abundant but becoming sparse). Biotope gradually transitional with previous biotope.	SS.SMx.CMx.ClloMx	SB:MX	-
D1-3.01	Maerl gravel (60%) and stone gravel (20%) with pebbles (5%) and shells (10%), with increasing sand content (5% overall); live maerl (<1%).	Saccharina latissima (C overall but A locally) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa. Patchy turf of foliose (R) and filiform (C) red algae including Bonnemaisonia asparagoides?, with brown algae including Desmarestia ligulata, D. aculeata and Dictyota dichotoma. Stones encrusted with pink coralline algae (R) and serpulid worms (P). Sediment with Cerianthus lloydii (C, locally A) and sparse live maerl thalli (R, <1%). Asterias rubens (P), Crossaster papposus (P), Marthasterias glacialis (F), Porania pulvillus (R), Gobiidae sp. (P).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-3.02	Mixed substrate of sand (55%, increasing along run), shell and stone gravel (30%), maerl gravel (5% but denser initially), pebbles (5%), shells (5%) and scattered cobbles (<1%) and boulders (<1%).	Stones and shells with hydroids (O) including Nemertesia antennina, Alcyonium digitatum (R), serpulid worms (P), solitary ascidians (O) including Corella parallelogramma and Ascidia virginea, and encrusting pink coralline algae (R), Ulva sp. (R), filiform red algae (O, initially F) and foliose red algae (R); Saccharina latissima (O, much probably drift), Desmarestia aculeata (R), D. ligulata (R); Saccharina fronds with Electra pilosa and Membranipora membranacea. Munida rugosa (R), Inachus sp. (P), Cancer pagurus (O), mollusc egg strings, Buccinum undatum (P), Pecten maximus (P), Aequipecten opercularis (F), Crossaster papposus (O), Marthasterias glacialis (O), Asterias rubens (O), Porania pulvillus (O), Luidia ciliaris (O), Henricia sp. (P), small teleost sp. (R). Sediment with small holes, worm casts, Chaetopterus variopedatus, Bonellidae sp. (P), bivalve siphons and Cerianthus lloydii (C). Biotope very gradually transitional with previous biotope and transitioning towards more homogeneous sandy sediment biotope at end.	SS.SMx.CMx.ClloMx	SB:MX	-
D1-4.01	Megaripples of maerl gravel (70%) and stone gravel (20%) with pebbles (<1%) and shells (10%) concentrated in troughs.	Shells and stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting tufts of filiform red algae (O). Saccharina latissima (O, probably largely drift material) with fronds encrusted with Membranipora membranacea and Electra pilosa, Desmarestia aculeata (R). Live maerl thalli (R, <1%), Cerianthus lloydii (A), Chaetopterus variopedatus (P), Liocarcinus sp.? (P), Marthasterias glacialis (P), Asterias rubens (F),	SS.SCS.ICS	SB:GS	-
D1-4.02	Maerl gravel (60%) and stone gravel (15%) with shells (10%) and pebbles (5%) on sand (10%); boulders (<1%).	Saccharina latissima (O) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa. Patchy turf of foliose red algae (R) including Delesseria sanguinea/Phycodrys rubens and filiform red algae (C) including Bonnemaisonia asparagoides?, with brown algae including Desmarestia spp.. Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting hydroid clumps (F), particularly Nemertesia antennina (F) and solitary ascidians (P). Sediment with Cerianthus lloydii (A), Chaetopterus variopedatus (P) and sparse live maerl thalli (R, <1%). Pecten maximus (P), Marthasterias glacialis (F), Porania pulvillus (O).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-4.03	Maerl gravel (60%) and stone gravel (15%) with shells (10%) and pebbles (5%) on sand (10%).	Poor visibility. Bed of Ophiocomina nigra (S) with shells and stones encrusted with pink coralline algae (R) and serpulid worms and supporting hydroids (F) including Nemertesia antennina (F) and filiform red algae (O). Marthasterias glacialis (P), Luidia ciliaris (P).	SS.SMx.CMx.OphMx	SB:MX	-
D1-4.04	Mixed sediment of maerl gravel (20% but higher initially), stone and shell gravel (30%), silty sand (45%, but higher in latter half), shells (5%) and boulders (<1%).	Stones and shells with hydroids (F) including Nemertesia antennina (F) and N. ramosa (P), serpulid worms (P), Balanus spp. (R), Omalosecosa ramulosa (R), Ascidia virginea (O), and encrusting pink coralline algae (R), Ulva sp. (R, possibly drift), filiform red algae (R) and foliose red algae (R); Saccharina latissima (O, much probably drift). Munida rugosa (O), Cancer pagurus (P), mollusc egg strings, Nudibranchia sp. (R), Pecten maximus (P), Aequipecten opercularis (O), Crossaster papposus (O), Marthasterias glacialis (P), Asterias rubens (F), Porania pulvillus (F), Luidia ciliaris (P), Henricia sp. (P), Echinus esculentus (O), small teleost sp. (R). Sediment with Chaetopterus variopedatus and Cerianthus lloydii (A, C towards end).	SS.SMx.CMx.ClloMx.Nem	SB:MX	-
D1-5.01	Maerl gravel (60%) and stone gravel (20%) with pebbles (5%) and shells (10%), sand (5% overall but increasing towards end); live maerl (<1%). Boulders and cobbles present but largely obscured by kelp.	Saccharina latissima (C overall but A locally over large area) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa. Patchy turf of foliose red algae (R) including Delesseria sanguinea/Phycodrys rubens, and filiform red algae (C) including Bonnemaisonia asparagoides?, with brown algae including Desmarestia spp. (F) including at least D. ligulata and D. aculeata, and Chorda filum (R). Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P). Sediment with Cerianthus lloydii (C, locally A), Chaetopterus variopedatus (P), Neopentadactyla mixta (P, 1 seen) and sparse live maerl thalli (R, <1%). Pecten maximus (P), Crossaster papposus (P), Marthasterias glacialis (F), Luidia ciliaris (F), Astropecten irregularis (P), Porania pulvillus (O), Gobiidae sp. (P).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D1-5.02	Silty sand (25%) with maerl gravel (45%), stone gravel (10%), pebbles (5%) and shells (15%); cobbles (<1%), boulders (<1%).	Stones and shells with clumps of hydroids (O) including Nemertesia ramosa, Alcyonium digitatum (R), serpulid worms (P), Balanus spp. (R), Omalosecosa ramulosa (R), solitary ascidians (R) and filiform red algae (O, some possibly drift), foliose red algae (R), pink encrusting coralline algae (R), Desmarestia spp. (R) and Saccharina latissima (O, possibly drift). Sediment with Cerianthus lloydii (A) and sparse live maerl thalli (R, <1%). Inachus sp. (P), Cancer pagurus (P), Necora puber (R), Liocarcinus sp. (R), Pecten maximus (P), Crossaster papposus (F), Marthasterias glacialis (P), Porania pulvillus (F), Luidia ciliaris (P), Echinus esculentus (P), small teleost sp. (R). Biotope gradually transitional with previous biotope.	SS.SMx.CMx.ClloMx	SB:MX	-
D1-6.01	Slightly mixed sediment of muddy sand (70%) with gravel (20%), pebbles (5%), shells (5%), cobbles (<1%) and boulders (<1%).	Stones and shells with clumps of hydroids (O) including Nemertesia antennina, Balanus spp. (R), Omalosecosa ramulosa (R), Ascidia virginea (R), Corella parallelogramma (F) and filiform (R) and foliose (R) red algae, some possibly drift; Saccharina latissima (R, probably all drift). Aphrodita aculeata (P), Munida rugosa (O), Paguridae spp. (R), Turritella communis (F locally), Buccinum undatum (R), mollusc egg string (P), Pecten maximus (R), Aequipecten opercularis (F), Crossaster papposus (P), Asterias rubens (O), Henricia spp. (R), Porania pulvillus (O), Luidia ciliaris (O), Ophiura ophiura (O, initially F), teleost sp. (R). Sediment with small holes and Cerianthus lloydii (C locally but generally sparse).	SS.SMx.CMx	-	-
D1-6.02	Cohesive muddy sand (98%) with shell gravel (1%), shells (1%) and cobbles (<1%).	Sediment with small holes, worm casts, Terebellidae spp. tentacles, emergent infaunal tubes and very sparse, small burrows. Hydroids (O) including Nemertesia ramosa and Halecium halecinum?, Munida rugosa (F), Paguridae sp. (R), Liocarcinus depurator (R), Turritella communis (F, at least locally), Buccinum undatum (R), Aequipecten opercularis (F), Marthasterias glacialis (P), Asterias rubens (O), Porania pulvillus (O), Astropecten irregularis? (R), Ophiura ophiura (F), teleost spp. (R), Corella parallelogramma (O), Callionymus spp. (R). Algal debris, although some filiform red algae probably functional (R). Areas of harder, transitional habitat at start and end.	SS.SMu.CSaMu	-	-
D1-6.03	Mixed substrate of muddy sand (75%) with shell gravel (14%), pebbles (5%), shells and large shell debris (5%), cobbles (1%) and boulders (<1%).	Stones and shells with clumps of hydroids (F) including Nemertesia antennina and N ramosa, serpulid worms (P), Omalosecosa ramulosa (F), Corella parallelogramma (O), Botryllus schlosseri? (R), and filiform red algae (R, some possibly drift). Munida rugosa (F), Paguridae sp. (R), Liocarcinus sp. (P), Aequipecten opercularis (P), Crossaster papposus (P), Porania pulvillus (O), Ophiura ophiura (P), Callionymus sp. (P). Sediment with small holes, Terebellidae spp. tentacles and emergent infaunal tubes.	SS.SMx.CMx	-	-
D1-6.04	Cohesive muddy sand (98%) with shell gravel (1%), shells (1%) and pebbles (<1%).	Sediment with small holes, Terebellidae spp. tentacles, emergent infaunal tubes and very sparse, small burrows. Stone and shell material supporting hydroids (F) including Nemertesia ramosa and N. antennina, and Omalosecosa ramulosa (F). Munida rugosa (F), Paguridae sp. (P), Liocarcinus sp. (P), Turritella communis (P), Nudibranchia sp. (P), Ophiura ophiura (F), teleost sp. (P), Corella parallelogramma (O), Callionymus spp. (O). Algal debris, although some filiform red algae probably functional (R).	SS.SMu.CSaMu	-	-
T1-6	Mud (100%).	Very poor visibility. Moderate density of megafaunal burrows including Calocaris macandreae (F-C) and Nephrops norvegicus (P, 8 animals seen). Caridea sp. (P), Asterias rubens (F). Euphausiids in water column.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-7.01	Mud (75%) with boulders (20%) and cobbles (5%).	Poor visibility. Silted stones supporting hydroid turf (A).	SS.SMu.CFiMu, CR.LCR	RF:ST	-
T1-7.02	Sandy mud (99%) with shell gravel (1%), boulders (<1%) and cobbles (<1%) and pebbles (<1%) towards end.	Stones with hydroids (where locally S but R overall). Sediment with emergent infaunal tubes, Virgularia mirabilis (R), Cerianthus lloydii (R) and lightly burrowed by megafauna, except for Nephrops norvegicus (C, 9 animals seen). Urticina sp. (R), Aphrodita aculeata (R), Munida rugosa (O), Caridea sp. (P), Pagurus bernhardus (R), Liocarcinus spp. (O) including L. depurator, Turritella communis shells (P, occupancy unknown, except for one with Paguridae sp.), Aporrhais pespelicani (P), Asterias rubens (O), Porania pulvillus (R), small teleost spp. (R), Lumpenus lampretaeformis (P).Glyptocephalus cyanoglossus? (O).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-7.03	Mud (50%) with boulders (45%, possibly including bedrock) and cobbles (5%).	Poor visibility. Silted stones supporting hydroid turf (A). Munida rugosa (F), Paguridae sp.? (P), Cancer pagurus (P). Trapped fishing net.	SS.SMu.CFiMu, CR.LCR	RF:ST	-
T1-7.04	Sandy mud (99%) with shell gravel (1%); scattered boulders (<1%) and cobbles (<1%) at end.	Stones with hydroids (where A but R overall). Sediment with emergent infaunal tubes and lightly burrowed by megafauna, except for Nephrops norvegicus (C, 3 animals seen). Munida rugosa (F),Turritella communis shells (P, occupancy unknown), Porania pulvillus? (P), small teleost sp. (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T1-7.05	Bedrock (100%).	Silted bedrock with hydroid turf (A). Asterias rubens (P).	CR.LCR	RF:BR	-
T1-8	Sandy mud (99%) with shell gravel (1%); sparse boulders (<1%) and cobbles (<1%).	Stones with hydroids (where locally C but R overall). Sediment with emergent infaunal tubes, low mounds, worm casts, Virgularia mirabilis (O), Funiculina quadrangularis (single colony), Sabella pavonina tubes? (R) and lightly burrowed by megafauna, except for Nephrops norvegicus (C, 18 animals seen); Calocaris macandreae (O). Munida rugosa (O), Paguridae spp. (O) including Pagurus bernhardus (R), Liocarcinus spp. (R), Turritella communis shells (P, occupancy unknown), Aequipecten opercularis (R), Asterias rubens (F), small teleost spp. (O), Lumpenus lampretaeformis (P).Glyptocephalus cyanoglossus? (F).	SS.SMu.CFiMu.SpnMeg	-	BM:SB, BM:FQ
T1-9	Mud (100%) becoming sandy mud with scattered cobbles, boulders and possibly bedrock towards end (all <1%).	Poor visibility and no photos. Mud moderately densely burrowed (diminishing towards end) by Calocaris macandreae (C, locally F) and Nephrops norvegicus (C, 9 animals seen) and supporting Virgularia mirabilis (R) and Cerianthus lloydii (R). Caridea sp. (P), Asterias rubens (P), teleost spp. (O), small Glyptocephalus cyanoglossus? (at least O). Dense euphausiids in water column towards end of run. Scattered stones and possibly bedrock toweards end of run with hydroids (where A); visibility too poor to determine whether separate biotope (CR.LCR).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
LBN-1.01	Maerl gravel (70%) and stone gravel (20%) with pebbles (5%) and shells (5%, but much greater at start and end).	Patchy, mostly short algal turf (A, locally C) dominated by filiform reds (C) and filiform browns (C) with Dictyota dichtoma (P) and mostly small Saccharina latissima (F), larger fronds with Membranipora membranacea; Ulva sp. (R). Live maerl reaches around 3% (R) locally but overall <1%. Cerianthus lloydii (C), Liocarcinus sp. (P), Asterias rubens (F), Marthasterias glacialis (C), Astropecten irregularis? (O), Luidia ciliaris (P), Henricia sp. (P), small teleost sp. (P). Pebbles encrusted with pink coralline algae (R).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
LBN-1.02	Dense shells (65%) on silty sand (20%) with gravel (15%).	Short turf of filiform/filamentous red algae (A) with Ulva sp. (R) and mostly small Saccharina latissima (F). Live maerl thalli (R, <1%). Marthasterias glacialis (P), Asterias rubens (F), Porania pulvillus (F), Henricia sp. (P), Ascidiella aspersa (P), small teleost spp. (F). Shells encrusted with pink coralline algae (O) and serpulid worms.	SS.SMp.KSwSS.LsacR	SB:KS	KS
LBN-1.03	Dense shells (40%) and live maerl (45%) with patches of silty sand (10%) and dead maerl (5%).	Rich maerl bed with live maerl overall around 45% (A) but denser locally. Shells encrusted with serpulid worms including Spirobranchus spp., and pink coralline algae (R). Short turf of mainly filamentous red algae (A) with filiform brown algae (F, at least locally) including Desmarestia spp., Chorda filum (F), and Saccharina latissima (F) supporting Membranipora membranacea and Electra pilosa, and Ulva sp. (R). Chaetopterus variopedatus (P), Liocarcinus spp. (O) including L. depurator (O), Necora puber (O), Cancer pagurus (P), Marthasterias glacialis (F), Crossaster papposus (F), Henricia spp. (O), Astropecten irregularis? (O), Porania pulvillus (F), Ascidiella aspersa (C locally), Scyliorhinus sp. (P), small teleost spp. (O) including Pholis gunnellus (P).	SS.SMp.Mrl.Pcal.R	SB:MB	MB
LBN-1.04	Silty sand (65%) with maerl gravel (30%) and shells (5%, locally greater).	Poor visibility and no photos. Patchy turf of filamentous red algae (C, locally A) with filiform brown algae (P), Chorda filum (C) and Saccharina latissima (F) with Membranipora membranacea. Sediment with Cerianthus lloydii (C, at least locall) and live maerl thalli (R, <1%). Marthasterias glacialis (F).	SS.SMp.KSwSS.LsacR	SB:KS	KS
LBN-1.05	Substrate largely obscured by algae but includes live maerl (possibly around 30%), shells (30%) with patches of maerl and stone gravel on muddy sand.	Maerl bed with density possibly around 30% (C), although much may be obscured by dense turf of filamentous red algae (S). Other algae include filiform browns (O), Asperococcus bullosus (R), Chorda filum (C), Saccharina latissima (F) with Membranipora membranacea, and Ulva sp. (R). Serpulid worms (P), Liocarcinus spp. (O) including L. depurator, Necora puber (P), Marthasterias glacialis (F), Porania pulvillus (O), Pholis gunnellus (P).	SS.SMp.Mrl.Pcal.R	SB:MB	MB
LBN-1.06	Dense shells (40%) on silty sand (45%) with maerl gravel (10%), pebbles (5%) and cobbles (<1%).	Turf of filamentous red algae (A, locally C) with filiform red (P) and brown algae (P), Saccharina latissima (F,locally C) with Membranipora membranacea and Electra pilosa, and Ulva sp. (R). Live maerl thalli present in places but apparently overall <1% (R). Chaetopterus variopedatus (P), Liocarcinus sp. (P), Pecten maximus (P), Archidoris pseudoargus? (O) and egg string, Marthasterias glacialis (F), Luidia ciliaris (P), Henricia spp. (O), Porania pulvillus (O), Ascidiella aspersa (F), small teleost spp. (O). Shells encrusted with serpulid worms and pink coralline algae (R).	SS.SMp.KSwSS.LsacR	SB:KS	KS
D3-1	Maerl (35%) and stone (30%) gravel with shells (10%) and pebbles (5%); patches dominated by sand (overall 20%).	Saccharina latissima (A, but locally C over large areas) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa; Chorda filum (C). Rich algal turf (C, locally A) of filiform and filamentous reds (C) including Bonnemaisonia asparagoides (C) and possibly Dasysiphonia japonica (P), foliose reds (R), brown algae including Dictyota dichotoma (P) and Desmarestia spp. (O) including at least D. aculeata, and Ulva sp. (R). Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P). Sediment with Cerianthus lloydii (P) and Chaetopterus variopedatus (P). Necora puber (P), Liocarcinus depurator (P), Marthasterias glacialis (C), Luidia ciliaris (F), Ascidia mentula (P), Ascidiella aspersa (P), small teleost sp. (P). Substrate varies regarding relative proportion of gravel and sand although much of it obscured by dense kelp. Interpreted as LsacR.Gv and LsacR.Sa biotopes present, although transitional areas widely present and impossible to clearly delineate the two biotopes.	SS.SMp.KSwSS.LsacR.Gv, SS.SMp.KSwSS.LsacR.Sa	SB:KS	KS
D3-2.01	Muddy sand (90%) with shell gravel (5%) and scattered shells (3%), pebbles (2%), cobbles (<1%) and boulders (<1%).	Sediment with small holes, low mounds, emergent infaunal tubes and sparse Cerianthus lloydii (O). Hydroids (O, locally F) including Nemertesia ramosa and N. antennina, Munida rugosa (F), Paguridae sp. (R), Liocarcinus sp. (R), Turritella communis (shells present, occupancy unknown), Aequipecten opercularis (O), Pecten maximus? (P), Asterias rubens? (P), Porania pulvillus (R), Luidia ciliaris (P), solitary ascidians (O) including Ascidiella aspersa (O), Gobiidae sp. (P). Algal debris, although some filiform red algae possibly functional (R).	SS.SSa.CMuSa	-	-
D3-2.02	Mixed substrate of muddy sand (60%) with gravel (10%), shells (5%), pebbles (15%), cobbles (10%) and boulders (<1%).	Stones supporting hydoids (F) including Nemertesia antennina (F) and N. ramosa (F), Alcyonidium diaphanum (R), Omalosecosa ramulosa (F, locally C) and solitary ascidians (F) including Corella parallelogramma, Ascidia virginea and A. mentula. Drift algae present with some filiform reds probably living (R). Sediment with Cerianthus lloydii (O), small holes, emergent infaunal tubes and some low mounds. Munida rugosa (F), Paguridae sp. (R), Necora puber (P), Liocarcinus spp. (O), Cancer pagurus (P), Buccinum undatum egg masses (P), Turritella communis shells (occupancy unknown), Pecten maximus (R), Aequipecten opercularis (O), Crossaster papposus (P), Asterias rubens (P), Porania pulvillus (F), Luidia ciliaris (O), Henricia spp. (O). Broad transitional areas at start and end of run.	SS.SMx.CMx.ClloMx.Nem	-	-
D3-2.03	Muddy sand (92%) with shell gravel (5%) and scattered shells (3%) and cobbles (<1%).	Sediment with small holes, worm casts, low mounds, emergent infaunal tubes and sparse Cerianthus lloydii (P). Hydroids (R) including Nemertesia antennina, Munida rugosa (F), Paguridae spp. (O), Liocarcinus sp. (P), Turritella communis (shells present, occupancy unknown), Aequipecten opercularis (O), Asterias rubens (P). Algal debris, although some filiform red algae possibly functional (R). Working creel at end of run.	SS.SSa.CMuSa	-	-
D3-3.01	Maerl (50%) and stone (30%) gravel with shells (15%) and pebbles (5%).	Saccharina latissima (C, locally A) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa; Chorda filum (C). Algal turf (C, locally A) of filiform and filamentous reds (C) including Bonnemaisonia asparagoides (C), brown algae including Desmarestia spp. (P), and Ulva sp. (R). Stones and shells encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P). Marthasterias glacialis (C), Asterias rubens (P), Ascidiella aspersa (A locally). Broad transitional area with following biotope.	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D3-3.02	Mixed substrate of silty sand (50%) with shell and stone gravel (20%), maerl gravel (2%, locally greater), shells (3%), pebbles (20%), cobbles (5%) and boulders (<1%).	Stones and shells encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydoids (O) including Nemertesia antennina and N. ramosa, solitary ascidians (locally A) including Ascidia virginea and Ascidiella aspersa (locally A), and algae including filiform reds (O), Desmarestia spp. (R), Ulva sp. (R) and Saccharina latissima (O) with Electra pilosa and Membranipora membranacea. Sediment with Cerianthus lloydii (C), emergent infaunal tubes and very sparse live maerl thalli (R, <1%). Munida rugosa (O), Liocarcinus sp. (P), Aequipecten opercularis (O), Marthasterias glacialis (P), Porania pulvillus (O), Astropecten irregularis (R), Henricia sp. (R), Scyliorhinus canicula (P).	SS.SMx.CMx.ClloMx	SB:MX	-
D3-3.03	Mixed substrate of silty sand (52%) with shell and stone gravel (20%), shells (2%), pebbles (20%), cobbles (5%) and boulders (1%).	Stones encrusted with pink coralline algae (R) and supporting hydoids (O) including Nemertesia antennina and N. ramosa, solitary ascidians (O) including Ascidia virginea and A. mentula, and algae including filiform reds (R) and Desmarestia sp.? (R); Saccharina latissima (P but drift). Sediment with worm casts. Munida rugosa (O), Marthasterias glacialis (P), Porania pulvillus (O), Luidia ciliaris (P), Henricia spp. (P).	SS.SMx.CMx	-	-
D3-3.04	Muddy sand with shell gravel (1%) and shells and large shell fragments (1%).	Sediment with small holes, sparse low mounds, worm casts and Cerianthus lloydii (O). Drift algae present but some filiform red algae possibly attached and functional (R). Nemertesia antennina (P), Munida rugosa (F), sparse Turritella communis shells (occupancy unknown), Porania pulvillus (P), Gobiidae sp. (P). No sharp boundary between this biotope of non-cohesive sediment and following biotope of cohesive sediment.	SS.SSa.CMuSa	-	-
D3-3.05	Muddy sand (95%) with scattered shells (5%, locally denser); boulders (<1%).	Shells supporting occasional hydroids including Nemertesia antennina and N. ramosa. Sediment with Cerianthus lloydii (P), small holes, worm casts, emergent infaunal tubes, low mounds and very sparse megafaunal burrows including Nephrops norvegicus. Aphrodita aculeata (R), Munida rugosa (F), Paguridae sp. (R), Turritella communis (F locally), Buccinum undatum (possibly dead), Aequipecten opercularis (R), Asterias rubens (F), Porania pulvillus (O). Part of depth-related continuum from non-cohesive to cohesive sediment and firm boundaries cannot be established.	SS.SMu.CSaMu	-	-
D3-3.06	Cohesive muddy sand or possibly sandy mud with shell gravel (1%), shells and large shell fragments (1%), and sparse silted boulders (<1%) and cobbles (<1%).	Sediment with emergent infaunal tubes, low mounds, worm casts and lightly burrowed by megafauna apart from Nephrops norvegicus (C, 4 animals seen). Stones with hydroids (O) including Nemertesia ramosa. Munida rugosa (F), Turritella communis (locally C), Asterias rubens (P), Porania pulvillus (R), teleost sp. (R), Callionymus sp. (R). Start boundary cannot be defined precisely.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D3-4	Mixed substrate varying in composition along run but overall silty sand (15%), maerl gravel (20%), stone gravel (20%), shell gravel (20%), shells (15%), pebbles (10%), cobbles (<1%), boulders (<1%).	Sediment with dense Cerianthus lloydii (C, locally A), Chaetopterus variopedatus? (P) and bivalve siphons (P). Stones and shells encrusted with pink coralline algae (R), serpulid worms and Balanus spp. including B. balanus, and supporting filiform red algae (R), Ulva sp. (R), hydroids (O) including Nemertesia antennina, Ascidiella aspersa (F, but A in first third) and Corella parallelogramma (O). Saccharina latissima (O) with Membranipora membranacea and Electra pilosa but probably drift material. Munida rugosa (R), Liocarcinus spp. (R) including L. depurator, Cancer pagurus (F), Pecten maximus (R), Aequipecten opercularis (R), Marthasterias glacialis (F, but C in first third), Asterias rubens (P), Astropecten irregularis (R), Luidia ciliaris (P), Henricia sp. (P), Porania pulvillus (O). Dense shoal of Ammodytidae sp.? at start of run. Large piece of discarded plastic sheeting.	SS.SMx.CMx.ClloMx	SB:MX	SE?
D3-5.01	Mixed substrate of silty sand (40%), shell gravel (30%), stone gravel (10%), shells and large shell material (5%), pebbles (10%), cobbles (5%) and boulders (<1%).	Stones encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydroids (F, locally A on boulders) including Nemertesia ramosa and N antennina, Omalosecosa ramulosa (R) and solitary ascidians (F, locally C) including Ascidiella aspersa and Ascidia virginea (F). Cerianthus lloydii (A), Munida rugosa (F), Liocarcinus sp.? (P), Aequipecten opercularis (F), Asterias rubens (C), Porania pulvillus (F), Henricia sp. (F), drift algae.	SS.SMx.CMx.ClloMx.Nem	-	-
D3-5.02	Mixed substrate of scattered pebbles (15%), cobbles (10%) and boulders (5%) on silty sand (30%) with shell gravel (25%), stone gravel (10%) and shells (5%).	Stones sparsely encrusted with pink coralline algae (R), serpulid worms and Balanus spp. (P) and supporting hydroids (F, A on boulders)) including Nemertesia ramosa (F) and N. antennina (F), mollusc egg strings, Omalosecosa ramulosa (R), Alcyonidium diaphanum (O), as well as solitary ascidians (C, locally A) including Ascidia virginea, A. mentula, Ascidiella aspersa (locally A) and Ciona intestinalis (locally A). Cerianthus lloydii (C), Cancer pagurus (P), Antedon spp. (F), Asterias rubens (C), Luidia ciliaris (F), Porania pulvillus (O), Marthasterias glacialis (P), Echinus esculentus (P). Considered a mosaic of AntAsH on boulders and CMx.ClloMx.Nem elsewhere.	SS.SMx.CMx.ClloMx.Nem, CR.LCR.BrAs.AntAsH	RF:ST	-
D3-5.03	Mixed substrate of silty sand (55%), gravel (25%), shells and large shell material (5%), pebbles (10%), cobbles (5%) and boulders (<1%).	Stones encrusted with pink coralline algae (R), serpulid worms (P) and Balanus spp. (P) and supporting hydroids (F) including Nemertesia ramosa (F) and N antennina (F), erect bryozoans (P) and solitary ascidians (F, A initially) including Ascidiella aspersa (A locally), Corella parallelogramma (P) and Ascidia mentula? (P). Cerianthus lloydii (C), Liocarcinus sp. (P), Buccinum undatum egg mass, Aequipecten opercularis (O), Pecten maximus? (P), Asterias rubens (F), Porania pulvillus (F), Echinus esculentus (P), drift algae.	SS.SMx.CMx.ClloMx.Nem	-	-
D3-5.04	Muddy sand (94%) with shell gravel (5%) and shells (1%); cobbles (<1%).	Sediment with Cerianthus lloydii (P), worm casts (P), emergent infaunal tubes (P) and small holes. Hydroids (F) including Nemertesia antennina (F), Munida rugosa (O), Turritella communis shells (occupancy unknown), Aequipecten opercularis (O), Solaster endeca (P). Gradual transition with following biotope.	SS.SSa.CMuSa	-	-
D3-5.05	Cohesive muddy sand (95%) with scattered shells (5%), cobbles (<1%) and boulders (<1%).	Sediment with Cerianthus lloydii (F), small holes, emergent infaunal tubes including possibly Chaetopterus variopedatus (P), worm casts and very sparse, small, megafaunal burrows; flattened and moribund Virgularia mirabilis (R). Shells and stones with hydroid clumps (O, but A on boulders) including Nemertesia ramosa and N. antennina, Omalosecosa ramulosa (R) and solitary ascidians (O) including Ascidia virginea (O). Munida rugosa (F), Paguridae spp. (O), Liocarcinus spp. (O), Cancer pagurus (P), Turritella communis shells (some containing pagurids), Pecten maximus (R), Aequipecten opercularis (O), Asterias rubens (P), Porania pulvillus (O), Callionymus sp. (R), Scophthalmus rhombus? (P), drift algae.	SS.SMu.CSaMu	-	-
D3-5.06	Mixed substrate of muddy sand (70%) with pebbles (5%), shell gravel (10%), shells and large shell material (15%), cobbles (<1%) and boulders (<1%).	Stones and shells with hydroids (O) and filiform red algae (R). Chaetopterus variopedatus? (P), Munida rugosa (F), Paguridae spp. (P) including occupant of Turritella communis shell.	SS.SMx.CMx	-	-
D3-5.07	Cohesive muddy sand (93%) with scattered shells (5%), shell gravel (2%) and cobbles (<1%).	Sediment with small holes, emergent infaunal tubes, worm casts, Terbellidae sp. tentacles and very sparse, small, megafaunal burrows. Shells and stones with hydroid clumps (F) including Nemertesia antennina (F), Omalosecosa ramulosa (R), Corella parallelogramma (O) and filiform red algae (R). Munida rugosa (O), Paguridae spp. (O), Turritella communis shells (some containing pagurids), Pecten maximus (P), Asterias rubens (F).	SS.SMu.CSaMu	-	-
D3-6	Sandy mud (97%) with shell gravel (2%), shells (1%) and sparse cobbles (<1%) and boulders (<1%).	Visibility poor. Shells and stones with hydroids (R overall) including Nemertesia ramosa, and Novocrania anomala (P). Sediment lightly burrowed by megafauna, except for Nephrops norvegicus (C, 3 animals seen). Munida rugosa (F), Turritela communis shells (occupancy unknown), Asterias rubens (F), Porania pulvillus (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D3-7.01	Maerl gravel (55%) and stone gravel (20%) with muddy sand (5%), pebbles (5%) and shells (15%).	Saccharina latissima (C, locally A) with older fronds supporting some dense mats of Membranipora membranacea and Electra pilosa; Chorda filum (C). Red algal turf (C, locally A) of filiform (C) and foliose (R) species including Bonnemaisonia asparagoides, brown algae including Desmarestia spp. (O-F) with D. aculeata (P), and Dictyota dichotoma (P), and Ulva sp. (R). Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P). Cerianthus lloydii (F-C), Liocarcinus spp. (P), bivalve siphons (P), Marthasterias glacialis (C, locally A), Asterias rubens (O), Porania pulvillus (R), Luidia ciliaris (O), Ascidiella aspersa (F-C), Corella parallelogramma (P), small teleost spp. (O), Gadidae sp. (P), live maerl thalli (R, <1%).	SS.SMp.KSwSS.LsacR.Gv	SB:KS	KS
D3-7.02	Mixed substrate of silty sand (25%) with maerl gravel (20%), stone gravel (10%), shell gravel (20%), shells (20%, locally greater) and pebbles (5%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms (C locally) including Spirobranchus spp. Dense Ascidiella aspersa (A) and Cerianthus lloydii (A). Drift algae present but some live including filiform reds (F), Desmarestia spp. (R), Dictyota dichotoma (R), Ulva sp. (R) and Saccharina latissima (F) with Electra pilosa and Membranipora membranacea. Paguridae sp. (P), Liocarcinus depurator (P), Marthasterias glacialis (F), Porania pulvillus (O), Luidia ciliaris (P), small teleosts spp. (O), sparse live maerl thalli (R, <1%).	SS.SMx.CMx.ClloMx	SB:MX	-
T5-1	Sandy mud (99%) with shell gravel (1%) and sparse boulders (<1%).	Mud lightly burrowed by megafauna including Nephrops norvegicus (F, 3 animals seen) and Lumpenus lampretaeformis (P) and supporting Virgularia mirabilis (O), Cereus pedunculatus (R), Cerianthus lloydii (R) and emergent infaunal tubes. Aphrodita aculeata (R), Caridea sp. (P), Munida spp. (O), Paguridae spp. (O) including Pagurus bernhardus, Neptunea antiqua (R), Aequipecten opercularis (R), teleost spp. (O), small Glyptocephalus cyanoglossus? (O), Pleuronectidae sp. (P). Euphausiids in water column. Scattered boulders with hydroids (where A, overall R). Possible trawl scars around 00:18:35.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T5-2	Sandy mud (99%) with shell gravel (1%) becoming softer mud towards end; cobbles (<1%), boulders (<1%).	Mud fairly lightly burrowed for much of run by megafauna including Nephrops norvegicus (C, 60 animals seen), Lumpenus lampretaeformis (O) and Calocaris macandreae (F, but increasing to C towards end as depth incrreases and mud softens) and supporting Virgularia mirabilis (O), emergent infaunal tubes, worm casts and small mounds. Aphrodita aculeata (R), Caridea sp. (O), Munida sp. (R), Paguridae spp. (O) including Pagurus bernhardus, Liocarcinus sp. (R), Buccinum undatum? (R), Turritella communis (locally F), Asterias rubens (O), teleost spp. (O), small Glyptocephalus cyanoglossus? (O). Euphausiids in water column, locally dense. Sparse cobbles and boulders with hydroids (where A, overall R).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T5-3.01	Sandy mud (99%) with shell gravel (1%) becoming harder towards end; cobbles (<1%), boulders (<1%).	Mud lightly burrowed for most of run by megafauna including Nephrops norvegicus (F, 34 animals seen, initially C), Lumpenus lampretaeformis (O) and probably Calocaris macandreae (P). However burrow density initially higher and then gradual transition to following biotope with very sparse burrows. Mud supports Virgularia mirabilis (O), Pennatula phosphorea (R), Cerianthus lloydii (R), emergent infaunal tubes, worm casts and small mounds. Aphrodita aculeata (R), Sabella pavonona tubes (R), Caridea sp. (R), Munida spp. (O), Paguridae spp. (R) including Pagurus bernhardus, Liocarcinus sp. (R), Buccinum undatum (R), Turritella communis (P), Arctica islandica siphons (P, 1 seen), Nudibranchia sp. (R), Asterias rubens (O), Porania pulvillus (R), teleost spp. (O), small Glyptocephalus cyanoglossus? (F). Sparse cobbles, boulders and unidentified substrates with hydroids including Nemertesia ramosa (O) and N. antennina (R), and Urticina sp.? (R).	SS.SMu.CFiMu.SpnMeg	-	BM:SB, AI
T5-3.02	Sandy mud (97%) with shell gravel (2%) and shells (1%); cobbles (<1%), boulders (<1%).	Sediment with small holes but very sparsely burrowed. Sediment with Virgularia mirabilis (F), Cereus pedunculatus (O), emergent infaunal tubes, worm casts and Arctica islandica siphons (one seen). Stones and shells with hydroids (O) including Nemertesia ramosa (O) and Urticina sp.? (R). Sabella pavonina tubes, Munida spp. (O), Paguridae sp. (R), Liocarcinus sp. (R), Buccinum undatum? (R), Turritella communis shells (P, occupancy unknown), Ophiura ophiura (O), mollusc egg strings, Asterias rubens (P), teleost spp. (O), Pleuronectiformes sp. (P), Glyptocephalus cyanoglossus? (O). Gradual transition with previous biotope.	SS.SMu.CSaMu	-	AI
T3-V01	Soft mud.	Moderate density of megafaunal burrows including Nephrops norvegicus (C, 124 animals seen) and smaller burrows including Calocaris macandreae (P). Virgularia mirabilis (R), Munida rugosa (R), Liocarcinus sp.? (R), teleost spp. (R) including Pleuronectiformes spp. (P). Sediment with parallel streaks visible at several points along run (00:33:49 - 00:34:15, 00:37:49 - 00:39:21, 00:40:28 - 00:40:53), presumably trawl scars. Possible c.20 mm diameter cable also possibly present in case of last set of scars.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T3-V02	Soft mud.	Moderate density of megafaunal burrows including Nephrops norvegicus (C, 128 animals seen) and smaller burrows including Calocaris macandreae (P). Hydroids (R), Pennatula phosphorea (O), Cerianthus lloydii (R), Munida rugosa (O), Liocarcinus spp.? (R), Turritella communis shells (P, occupancy unknown), Asterias rubens (O), teleost spp. (O), Pleuronectiformes spp. (O), sparse emergent infaunal tubes.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T3-V03.01	Sandy mud	Light to moderate density of megafaunal burrows including Nephrops norvegicus (C, 2 animals seen) and smaller burrows including probably Calocaris macandreae (P). Sediment also supports dense Turritella communis (C) and with emergent infaunal tubes, small mounds, bivalve siphons and worm casts. Hydroids (R) including Nemertesia antennina, Pennatula phosphorea (R), Cerianthus lloydii (R), Myxicola infundibulum (P), Munida rugosa (O), Liocarcinus spp. (O), Asterias rubens (O), Pleuronectiformes sp. (P). Biotope transitional between SpnMeg and CSaMu and could be ascribed to either.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T3-V03.02	Sandy mud or muddy sand (80%) with scattered gravel (4%), pebbles (15%), cobbles (1%) and boulders (<1%).	Stones support hydroid turf (F). Munida rugosa (F), Cancer pagurus (P), Liocarcinus sp.? (P), Turritella communis shells (P, occupancy unknown).	SS.SMx.CMx	-	-
T3-V03.03	Soft mud with patches of harder, possibly sandier, mud.	Much of run of soft mud with moderate density of megafaunal burrowers including Nephrops norvegicus (C, 10 animals seen) and smaller species including probably Calocaris macandreae (P). Harder patches with slightly reduced burrow density but with Turritella communis (locally C). Many shells apparently occupied by mollusc, but some with Paguridae spp. (P). Cerianthus lloydii (P), Munida rugosa (O), Liocarcinus spp. (O), Goneplax rhomboides (P), Cancer pagurus (P), Asterias rubens (F), teleost spp. (O). Sediment with parallel streaks visible between 00:36:52 and 00:38:03, presumably trawl scars. Also very fine, dense, faint striations discernible in direction of travel for much of run, possibly of natural origin.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T3-V03.04	Sandy mud or muddy sand (75%) with scattered gravel (2%), pebbles (10%), cobbles (10%) and boulders (3%).	Stones support hydroid turf (F). Munida rugosa (P), Turritella communis shells (P, occupancy unknown), sparse, small megafaunal burrows.	SS.SMx.CMx	-	-
T3-V03.05	Sandy mud	Light to moderate density of megafaunal burrows including Nephrops norvegicus (C, 2 animals seen) and smaller burrows. Sediment also supports Turritella communis (C locally). Pennatula phosphorea (R), Liocarcinus spp. (O), Pecten maximus (P), Asterias rubens (P), teleost sp. (R). Fishing cable or rope at 00:56:19. In addition to Turritella tracks, sediment with fine, quasi-parallel striations for most of run in direction of travel - possibly of natural origin. Biotope transitional between SpnMeg and CSaMu.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T3-V03.06	Sandy mud or muddy sand (60%) with scattered gravel (5%), pebbles (20%), cobbles (10%) and boulders (5%).	Stones support hydroid turf (C) including Nemertesia antennina. Munida rugosa (P), Turritella communis shells (P, occupancy unknown), Asterias rubens (P), sparse, small megafaunal burrows.	SS.SMx.CMx	-	-
T3-V04	Soft mud with patches of harder, possibly sandier, mud; isolated boulders (<1%)..	Much of run of soft mud with moderate density of megafaunal burrowers including Nephrops norvegicus (C, 21 animals seen) and smaller species including Calocaris macandreae (P). Harder patches with slightly reduced burrow density but with Turritella communis (locally C). Hydroids (R) including Nemertesia antennina, Pennatula phosphorea (R), Cerianthus lloydii (R), Myxicola infundibulum (P), Munida rugosa (O), Liocarcinus depurator (R), Goneplax rhomboides (P), Cancer pagurus (P), Aequipecten opercularis (R), Asterias rubens (O), teleost spp. indet. (R), Callionymus spp. (R), Lampenus lampretaeformis (P). Discarded creel.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D4-V01.01	Slightly muddy fine sand (94%) with broken shell (5%) and whole shells (1%); sparsely scattered cobbles (<1%).	Sand punctured by small infaunal holes and supporting fairly sparse emergent tubes (O, possibly Cerianthus lloydii). Shell and stones with sparse hydroids (R) and Omalosecosa ramulosa (R). Munida rugosa (R), Pecten maximus? (R), Porania pulvillus (R), small Asteroidea sp. (R).	SS.SSa.CMuSa	-	-
D4-V01.02	Mixed substrate of scattered pebbles (20%) and shelly gravel (10%) on slightly muddy fine sand (25%) with silted cobbles (30%) and boulders (15%) but 100% in patches.	Stones sparsely encrusted with pink coralline algae (R) and Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F) including Nemertesia ramosa, Caryophyllia smithii (R) and Omalosecosa ramulosa (O), as well as solitary ascidians (O) including Ascidia virginea and A. mentula. Munida rugosa (F), Cancer pagurus (P), Pecten maximus (O), Antedon sp. (R), Asterias rubens (F), Luidia ciliaris (F), Porania pulvillus (O), Marthasterias glacialis (P), Echinus esculentus (F), Labrus mixtus (P). Sediment with sparse infaunal tubes (P). Considered a mosaic of AntAsH in boulder/cobble patches and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs.AntAsH	RF:ST	-
D4-V01.03	Slightly muddy fine sand (89%) with broken shell (10%) and whole shells (1%); sparsely scattered cobbles (<1%) and boulders (<1%).	Sand with sparse emergent tubes (R). Shell and stones with hydroid clumps (R). Munida rugosa (P), Porania pulvillus (O), small Asterias rubens? (F), Callionymus sp.? (P).	SS.SSa.CMuSa	-	-
D4-V01.04	Mixed substrate varying along run but mainly a gravelly (37%), slightly muddy sand (42%) with scattered pebbles (15%), shells (1%), cobbles (5%) and boulders (<1%).	Stones encrusted with Balanus spp. (R) and supporting hydroids (R) including Nemertesia ramosa?, Omalosecosa ramulosa (R) and Ascidia mentula (O). Munida rugosa (F), Liocarcinus sp. (O), Aequipecten opercularis (R), Luidia ciliaris (F), Porania pulvillus (O). Sediment with emergent tubes (P) and small holes and mounds.	SS.SMx.CMx	-	-
D4-V01.05	Mixed substrate varying along run but mainly a gravelly (10%), slightly muddy sand (40%) with pebbles (19%), shells (1%). Cobbles (25%) and boulders (5%) but denser in patches.	Stones sparsely encrusted with pink coralline algae (R), serpulid worms (P) and Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F, locally O) including Nemertesia ramosa (O), and Omalosecosa ramulosa (R), as well as Ascidia mentula (O). Munida rugosa (F), Liocarcinus sp. (R), Cancer pagurus (O), Necora puber? (R), Pecten maximus (O), Asterias rubens (F), Stichastrella rosea (R), Luidia ciliaris (O), Porania pulvillus (O), Marthasterias glacialis (P), teleost spp. (O) including Callionymus sp.? (R). Sediment with holes and infaunal tubes. Considered a mosaic of AntAsH in boulder/cobble patches and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs.AntAsH	RF:ST	-
D4-V01.06	Slightly muddy fine sand (80%) with broken shell (10%), stone gravel (2%) and whole shells (1%); pebbles (5%), cobbles (1%) and boulders (<1%).	Sand with holes and sparse emergent tubes (R). Shell and stones with hydroid clumps (R) and Omalosecosa ramulosa (R). Munida rugosa (O), Liocarcinus sp. (R), Pecten maximus (O), Porania pulvillus (O), Astropecten irregularis (R), Callionymus sp. (O).	SS.SSa.CMuSa	-	-
D4-V01.07	Mixed substrate varying along run but mainly a gravelly (35%), slightly muddy sand (44%) with scattered pebbles (15%), shells (1%), cobbles (5%) and boulders (<1%).	Stones encrusted with serpulid worms (P) and pink coralline algae (R) and supporting hydroids (O, locally F) and Omalosecosa ramulosa (R). Munida rugosa (F), Cancer pagurus (P), Pecten maximus (R), Marthasterias glacialis (P), Asterias rubens (F), Luidia ciliaris (F), Porania pulvillus (O), teleost sp. (R). Sediment with emergent tubes (P) and small holes and mounds.	SS.SMx.CMx	-	-
D4-V01.08	Largely dense, silted boulders (45%) and cobbles (30%), with infill of gravelly (5%), slightly muddy sand (10%) with pebbles (10%).	Stones sparsely encrusted with Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F), as well as Ascidia mentula (F). Munida rugosa (O), Cancer pagurus (F), Calliostoma zizyphinum (P), Antedon sp. (O), Asterias rubens (C), Stichastrella rosea? (O), Porania pulvillus (O), Marthasterias glacialis (F), Echinus esculentus (C), teleost spp. (O) including Labrus mixtus (P).	CR.LCR.BrAs.AntAsH	RF:ST	-
D4-V01.09	Slightly muddy sand (65%) with gravel (20%), pebbles (10%) and silted cobbles (5%) and boulders (<1%).	Stones supporting hydroids (O) and Omalosecosa ramulosa (R). Munida rugosa (F), Pecten maximus (P), Porania pulvillus (F). Sediment with emergent tubes (P) and small holes and mounds.	SS.SMx.CMx	-	-
D4-V01.10	Slightly muddy fine sand (85%) with gravel (10%), pebbles (4%), cobbles (1%) and boulders (<1%).	Sand with holes. Stones with hydroid clumps (R). Munida rugosa (O).	SS.SSa.CMuSa	-	-
D4-V01.11	Largely dense, silted boulders (35%) and cobbles (40%), with infill of gravelly (5%), slightly muddy sand (10%) with pebbles (10%).	Stones sparsely encrusted with Parasmittina trispinosa (R) and supporting turf of hydroids (F), Caryophyllia smithii (P), Omalosecosa ramulosa (R), Ascidia mentula (F) and A. virginea? (P). Munida rugosa (F), Asterias rubens (C), Marthasterias glacialis (C), Luidia ciliaris (P), Echinus esculentus (C).	CR.LCR.BrAs.AntAsH	RF:ST	-
D4-V01.12	Slightly muddy sand (65%) with broken shell (5%), stone gravel (15%), pebbles (5%) and silted cobbles (10%) and boulders (<1%).	Stones supporting hydroids (O) and Omalosecosa ramulosa (R). Marthasterias glacialis (P). Sediment with small mounds.	SS.SMx.CMx	-	-
D4-V01.13	Slightly muddy fine sand (94%) with broken shell (5%) and whole shells (1%); sparsely scattered pebbles (<1%) and cobbles (<1%).	Sand with small mounds and punctured by small infaunal holes and supporting possibly small Cerianthus lloydii (R). Shell and stones with sparse hydroids (R) and Omalosecosa ramulosa (R). Munida rugosa (O), Liocarcinus spp. (O), Pecten maximus (R), Crossaster papposus (P), Asterias rubens (P), Porania pulvillus (O), Marthasterias glacialis (P), Luidia ciliaris (O), Callionymus sp. (R), teleost sp. (R).	SS.SSa.CMuSa	-	-
D4-V02.01	Mixed sediment of silty sand (63%) and shell and stone gravel (30%) with proportions varying locally; pebbles (5%), shells (1%), cobbles (1%) and boulders (<1%).	Sediment with occasional small mounds, as well as small holes and emergent infaunal tubes; Small Cerianthus lloydii possibly present. Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting sparse hydroid clumps (R). Pecten maximus (R), Crossaster papposus (P), Marthasterias glacialis (O), Asterias rubens (O), Henricia sp. (R), Astropecten irregularis (R), Porania pulvillus (O).	SS.SMx.CMx	-	-
D4-V02.02	Mixed substrate of gravel (30%) and silty sand (20%) with pebbles (10%); patches of silted cobbles (25%) and boulders (15%), denser locally.	Stones encrusted with pink coralline algae (R) and Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F), as well as Ascidia mentula (F), foliose red algae (R) and filamentous/filiform red algae (R). Munida rugosa (F), Brachyura sp. (P), Asterias rubens (F), Crossaster papposus (F), Luidia ciliaris (P), Henricia sp. (O), Porania pulvillus (O), Marthasterias glacialis (F), Echinus esculentus (F), teleost spp. (O) including Labrus mixtus (O). Considered a mosaic of BrAs in boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V02.03	Mixed sediment of silty sand (50%) with shell and stone gravel (35%), shells (<1%), maerl gravel (5%) and pebbles (10%).	Sediment with small holes and emergent infaunal tubes (O). Stones and shells with serpulid worms (P), hydroids (R) and pink coralline algae (R). Pecten maximus (O), Porania pulvillus (O).	SS.SMx.CMx	-	-
D4-V02.04	Mixed substrate of gravel (25%) and silty sand (30%) with pebbles (4%) and shells (1%); patches of silted cobbles (10%) and boulders (30%), denser locally.	Stones encrusted with pink coralline algae (R) and Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F), as well as Ascidia mentula (O overall, F on boulders) and foliose red algae (R). Munida rugosa (F), Necora puber (O), Cancer pagurus (P), Liocarcinus sp. (P), Asterias rubens (C), Porania pulvillus (O), Marthasterias glacialis (F), teleost spp. (F) including Labrus mixtus (P) and Callionymus sp. (P). Sediment with small holes and emergent infaunal tubes, possibly Chaetopterus variopedatus (P). Considered a mosaic of BrAs in boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V02.05	Mixed sediment of silty sand (60%) with shell and stone gravel (35%), shells (1%), maerl gravel (1%) and pebbles (3%); cobbles and boulders (<1%).	Sediment with small mounds, as well as small holes and emergent infaunal tubes. Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting sparse hydroid clumps (R). Munida rugosa (R), Inachus sp. (R), Liocarcinus sp. (R), Pecten maximus (R), Marthasterias glacialis (F), Asterias rubens (F), Porania pulvillus (O).	SS.SMx.CMx	-	-
D4-V02.06	Mixed substrate of gravel (15%) and silty sand (30%) with pebbles (10%); patches of silted cobbles (30%) and boulders (15%), denser locally.	Stones encrusted with pink coralline algae (R), serpulid worms (P) and Parasmittina trispinosa (R) and supporting patchy turf of hydroids (F), as well as Ascidia mentula (O overall, F on boulders and cobbles), Caryophyllia smithii (R), foliose red algae (R) and filamentous/filiform red algae (R). Munida rugosa (O), Necora puber? (R), Cancer pagurus (P), Pecten maximus (O), Crossaster papposus (F), Asterias rubens (F), Porania pulvillus (O), Marthasterias glacialis (F), Luidia ciliaris (F), Echinus esculentus (F), teleost spp. (O) including Labrus mixtus (P). Sediment with small holes and emergent infaunal tubes. Considered a mosaic of BrAs in boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V02.07	Mixed sediment of silty sand (60%) with shell and stone gravel (30%), maerl gravel (5%), shells (1%), pebbles (4%); cobbles and boulders (<1%).	Sediment with small mounds, as well as small holes and emergent infaunal tubes. Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting sparse hydroid clumps (R). Cancer pagurus (P), Pecten maximus (R), Marthasterias glacialis (F), Asterias rubens (F), Luidia ciliaris (P), Astropoecten irregularis (R), Porania pulvillus (O).	SS.SMx.CMx	-	-
D4-V03	Cohesive muddy sand with shell gravel (4%), shells (1%) and sparse cobbles (<1%) and boulders (<1%).	Sediment with small holes, some paired and probably bivalve siphons, worm casts, emergent infaunal tubes, Cerianthus lloydii (R), Myxicola infundibulum (P) and sparse, mostly small, megafaunal burrows; also sediment mounds up to around 15 cm diameter, possibly those of Callianassa subterranea. Shells and stones support hydroid clumps (R). Munida rugosa (R), Liocarcinus spp. (O), Cancer pagurus (O), Buccinum undatum (R), Turritella communis (O), Pecten maximus (R), Asterias rubens (O), Luidia ciliaris (O), Marthasterias glacialis (O), Porania pulvillus (F), Astropecten irregularis (R), Ophiura sp. (R), Callionymus sp. (O), teleost spp. indet. (R). Many empty shells of Arctica islandica.	SS.SMu.CSaMu	-	-
T2-V01	Soft mud.	Moderate density of megafaunal burrows including Nephrops norvegicus (C, 18 animals seen) and smaller burrows including Calocaris macandreae (P). Pennatula phosphorea (O), Liocarcinus spp. (R) including L. depurator, Turritella communis (P), Porania pulvillus (R), Asterias rubens (O), teleost spp. (O) including Pleuronectiformes spp. (O), emergent infaunal tubes (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T4-V01	Soft mud.	Mud densely burrowed, particularly by Calocaris macandreae and Nephrops norvegicus (C, 83 animals seen). Calocaris density is difficult to determine due to operlapping clusters of burrows and burrows of other species but is at least C overall and A at least locally. Virgularia mirabilis (R), Pennatula phosphorea? (R), Aphrodita aculeata (R), Caridea sp. (R), Buccinum undatum (R), Asterias rubens (O), teleost spp. (R). Faint parallel scars at around 00:56:16 possibly representing old trawling disturbance.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T4-V02	Soft mud.	Mud moderately densely burrowed, particularly by Calocaris macandreae (C, locally F) and Nephrops norvegicus (C, 49 animals seen). Virgularia mirabilis (R), Metridium dianthus (P) on cobble, Caridea sp. (O), Asterias rubens (F), teleost spp. (O). Visibility latterly poor due to recent trawling activity in region.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T4-V03	Soft mud.	Mud moderately densely burrowed, particularly by Calocaris macandreae (C, locally F and possibly A) and Nephrops norvegicus (C, 41 animals seen). Urticina sp.? (R), Caridea sp. (R), Asterias rubens (F), teleost spp. (R). Visibility latterly poor due to recent trawling activity in region. Discarded plastic sack. Probable trawl scars visible intermittently (e.g. 00:03:30 - 00:04:30, 00:08:33 - 00:08:57).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D6-V01.01	Slightly muddy fine sand with shell gravel (5%) and shells (1%).	Sediment with small mounds, worm casts, emergent infaunal tubes and Cerianthus lloydii (R). Shells support serpulid worms (P) and clumps of hydroids (R), Flustra foliacea (R) and filiform red algae (R). Urticina felina (R), Munida rugosa (O), Liocarcinus spp. (R), Cancer pagurus (P), Turritella communis (F, at least locally), Aequipecten opercularis (R), Luidia ciliaris (O), Porania pulvillus (O), Ophiura ophiura (O, locally F), Callionymus sp. (R).	SS.SSa.CMuSa	-	-
D6-V01.02	Sand (50%) with gravel (15%), pebbles (20%), cobbles (15%) and shells (<1%).	Sediment with Cerianthus lloydii (C, at least locally). Stones support serpulid worms (P), Ascidia virginea? (P) and clumps of hydroids (O), bryozoans (R) and filiform red (O) and foliose red (R) algae. Urticina sp. (P), Munida rugosa (F), Liocarcinus sp. (P), Turritella communis shells (P, occupancy unknown), Henricia sp. (P), Porania pulvillus (F).	SS.SMx.CMx.ClloMx	-	-
D6-V02	Highly variable substrate but mostly mixed, fine-coarse sand (65%) with varying quantities of gravel (20%), pebbles (10%), shells (1%), cobbles (4%) and boulders (<1%).	Fauna visually dominated by Cerianthis lloydii (C over much of run, A locally); worm casts (P). Stones and shells encrusted with pink coralline algae (R) and serpulid worms and supporting hydroid clumps (O) including Nemertesia antennina, Alcyonium digitatum (R), Sagartia elegans? (R), Urticina spp. (R), Metridium dianthus (O), Omalosecosa ramulosa? (R), Flustra foliacea (R) and solitary ascidians (R) including Ascidia virginea and A. mentula. Munida rugosa (R), Liocarcinus spp. (R), Cancer pagurus (O), Necora puber (R), Turritella communis shells (P, occupancy unknown), Pecten maximus (R), Aequipecten opercularis (R), Marthasterias glacialis (O), Crossaster papposus (O), Asterias rubens (O), Porania pulvillus (F), Henricia sp. (R), Luidia ciliaris (O), Astropecten irregularis (R), Echinus esculentus (O), teleost spp. (O) including Callionymus spp. (P) and Pleuronectiformes sp. (P).	SS.SMx.CMx.ClloMx	-	-
D6-V03.01	Mixed sediment of sand (35%), stone gravel (35%), maerl gravel (20%) and shells (10%).	Fauna strongly visually dominated by Cerianthis lloydii (A). Shells encrusted with pink coralline algae (R) and serpulid worms and supporting Ascidiells aspersa (O) and filiform/filamentous red algae (R). Inachus sp. (P), Liocarcinus depurator (P), Cancer pagurus (P), Pecten maximus (O), Marthasterias glacialis (F), Asterias rubens (P), Porania pulvillus (F), Luidia ciliaris (P), Astropecten irregularis? (P), Echinus esculentus (P), teleost sp. (P), Pleuronectiformes sp. (P).	SS.SMx.CMx.ClloMx	SB:MX	-
D6-V03.02	Coarse sediment formed into megaripples in places and composed of sand (10%), stone gravel (40%), maerl gravel (35%), live maerl (<1%) and shells (15%).	Cerianthus lloydii (C, locally A), live maerl thalli (R, <1%). Shells and gravel encrusted with pink coralline algae (O) and serpulid worms (P) and supporting filamentous/filiform red algae (O) and foliose red algae (R), hydroids (R) and Ascidiella aspersa (P); Saccharina latissima (F, but apparently largely drift material). Inachus sp. (P), Liocarcinus spp. (R), Cancer pagurus (O), Pecten maximus (R), Marthasterias glacialis (F), Crossaster papposus (O), Asterias rubens (F), Porania pulvillus (F), Luidia ciliaris (P), Astropecten irregularis (R).	SS.SCS.CCS	SB:GS	-
D6-V03.03	Megaripples of maerl gravel (50%), live maerl (10%), stone gravel (20%), sand (5%) and shells (15%).	Shells and live maerl thalli (10%, F) tending to be concentrated in megaripple troughs. Other algae include encrusting pink corallines on shells and gravel (O), filamentous/filiform reds (O), foliose reds (R), Desmarestia aculeata (R) and Saccharina latissima (F, but probably largely drift material). Cerianthus lloydii (P), serpulid worms on shells (P), Liocarcinus depurator (R), Cancer pagurus (O), Marthasterias glacialis (F), Asterias rubens (O), Porania pulvillus (R), Luidia ciliaris (O), Astropecten irregularis (R), Ascidiella aspersa (A in patches), small teleost spp. (P).	SS.SMp.Mrl	SB:MB	MB
D6-V03.04	Coarse sediment formed into megaripples in places and composed of coarse sand/finely comminuted dead maerl (50%), maerl gravel (20%), shell and stone gravel (15%), live maerl (<1%) and shells (15%).	Cerianthus lloydii (C, locally A), live maerl thalli (R, <1%). Shells and gravel encrusted with pink coralline algae (O), red algae (R) and serpulid worms (P) and supporting filamentous/filiform red algae (O) including Scinaia interrupta (R) and foliose red algae (R), and Ascidiella aspersa (P); Saccharina latissima (P, but apparently largely drift material), Desmarestia aculeata (R). Cancer pagurus (F), Marthasterias glacialis (F), Asterias rubens (P), Luidia ciliaris (P).	SS.SCS.CCS	SB:GS	-
D6-V03.05	Mixed substrate of sand (50%) with shell and stone gravel (20%), maerl gravel (10%), pebbles (10%), shells (5%) and scattered boulders (3%) and cobbles (2%).	Boulders, cobbles and smaller stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting patchy algal turf of predominantly filiform/filamentous reds (C) with foliose reds (R) and Saccharina latissima (O), as well as hydroids (R) and dense Ascidiella aspersa (locally S). Cerianthus lloydii (C, locally A), Liocarcinus spp. (R), Cancer pagurus (F), Eledone cirrhosa? (P), Antedon sp. (R), Crossaster papposus (O), Marthasterias glacialis (F), Asterias rubens (P), Luidia ciliaris (F), Porania pulvillus (O), Echinus esculentus (P), small teleost sp. (R).	SS.SMp.KSwSS	SB:KS	KS
D6-V04.01	Sand (60%) with shell and stone gravel (30%), pebbles (5%), shells (5%) and scattered cobbles (<1%) and boulders (<1%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P) and supporting hydroids (O) including Nemertesia antennina, Urticina spp. (R), Flustra foliacea (R), and filiform/filamentous red algae (R, locally O). Cerianthus lloydii (C, locally A), Munida rugosa (O), Pagurus bernhardus (R), Cancer pagurus (O), Liocarcinus spp. (R), Buccinum undatum (P), Pecten maximus (R), Aequipecten opercularis (R), Marthasterias glacialis (P), Asterias rubens (P), Luidia ciliaris (O), Porania pulvillus (F), Ophiura ophiura (P), Echinus esculentus (P), small teleost spp. (R).	SS.SMx.CMx.ClloMx	-	-
D6-V04.02	Sand (50%) with shell and stone gravel (40%), pebbles (5%), shells (3%) and cobbles (2%).	Stones and shells encrusted with serpulid worms and supporting hydroids (R). Cerianthus lloydii (A), Ophiocomina nigra (A), Porania pulvillus (F).	SS.SMx.CMx.OphMx	-	-
D6-V04.03	Sand (35%) with shell and stone gravel (45%), pebbles (15%), shells (3%) and scattered cobbles (2%) and boulders (<1%).	Stones and shells encrusted with serpulid worms (P) and Parasmittina trispinosa? (R) and supporting hydroids (O) including Nemertesia antennina and N. ramosa, Flustra foliacea (R), solitary ascidians (R) including Ascidia virginea, and filiform/filamentous red algae (R). Cerianthus lloydii (A), Munida rugosa (O), Cancer pagurus (F), Pecten maximus (O), Marthasterias glacialis (P), Asterias rubens (F), Luidia ciliaris (F), Porania pulvillus (F), Ophiocomina nigra (R), small teleost sp. (R).	SS.SMx.CMx.ClloMx	-	-
D6-V04.04	Sand (35%) with shell and stone gravel (45%), pebbles (15%), shells (2%) and scattered cobbles (2%) and boulders (1%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms and supporting hydroids (O) including Nemertesia ramosa, and solitary ascidians (P). Cerianthus lloydii (A), Ophiocomina nigra (A), Luidia ciliaris (P), Porania pulvillus (O), small teleost spp. (F).	SS.SMx.CMx.OphMx	-	-
D6-V04.05	Sand (35%) with shell and stone gravel (45%), pebbles (10%), shells (3%) and scattered cobbles (2%) and boulders (5%).	Stones and shells encrusted with pink coralline algae (R) and serpulid worms (P) and supporting hydroids (O) including Nemertesia ramosa and Abietinaria abietina, Urticina sp. (P), Flustra foliacea (R) and solitary ascidians (P). Cerianthus lloydii (A), Cancer pagurus (P, Aequipecten opercularis (P), Asterias rubens (P), Luidia ciliaris (C), Porania pulvillus (F), small teleost spp. (F).	SS.SMx.CMx.ClloMx	-	-
D6-V04.06	Sand (45%) with shell and stone gravel (25%), pebbles (15%), shells (5%) and scattered cobbles (5%) and boulders (5%).	Stones encrusted with serpulid worms and supporting hydroids (O), Omalosecosa ramulosa (R) and solitary ascidians (A locally) including Ascidia virginea. Virgularia mirabilis (P), Cerianthus lloydii (A), Chaetopterus variopedatus? (P), Pecten maximus (P), Ophiocomina nigra (A, locally S), Luidia ciliaris (F), Henricia sp. (P), Porania pulvillus (F), small teleost sp. (P).	SS.SMx.CMx.OphMx	-	-
D6-V04.07	Sand (45%) with shell and stone gravel (35%), pebbles (10%), shells (3%) and scattered cobbles (5%) and boulders (2%).	Stones and shells encrusted with pink coralline algae (R), Parasmittina trispinosa? (R) and serpulid worms (P) and supporting hydroids (O) including Nemertesia antennina, Urticina sp. (P) and solitary ascidians (P). Cerianthus lloydii (A), Munida rugosa (O), Luidia ciliaris (F), Porania pulvillus (F), Ophiocomina nigra (F), small teleost sp. (P).	SS.SMx.CMx.ClloMx	-	-
D6-V04.08	Sand (55%) with shell and stone gravel (20%), pebbles (15%), shells (3%) and scattered cobbles (5%) and boulders (2%).	Stones encrusted with serpulid worms and pink coralline algae (R) and supporting hydroids (O). Cerianthus lloydii (A), Pecten maximus (O), Ophiocomina nigra (A), Asterias rubens (F), Luidia ciliaris (P), Porania pulvillus (F), Echinus esculentus (F), small teleost sp. (P).	SS.SMx.CMx.OphMx	-	-
D6-V04.09	Sand (50%) with shell and stone gravel (25%), pebbles (10%), shells (2%) and scattered cobbles (5%) and boulders (8%).	Stones and shells encrusted with pink coralline algae (R), Parasmittina trispinosa? (R) and serpulid worms (P) and supporting hydroids (O) including Nemertesia antennina and Abietinaria abietina, Flustra foliacea (R), Omalosecosa ramulosa (R) and solitary ascidians (O). Cerianthus lloydii (A), Munida rugosa (O), Cancer pagurus (P), Asterias rubens (P), Luidia ciliaris (P), Porania pulvillus (F), Ophiocomina nigra (O), small teleost spp. (F) including Labrus mixtus (P).	SS.SMx.CMx.ClloMx	-	-
D6-V04.10	Sand (55%) with shell and stone gravel (15%), pebbles (5%), shells (3%) and scattered cobbles (7%) and boulders (15%).	Stones encrusted with serpulid worms and Parasmittina trispinosa? (R) and supporting hydroids (O) including Nemertesia antennina, Urticina sp. (P), Bolocera tuediae (P), Flustra foliacea (R), Omalosecosa ramulosa (R), and solitary ascidians (O) including Ascidia virginea and A. mentula. Cerianthus lloydii (A), Munida rugosa (O), Cancer pagurus (P), Pecten maximus (O), Ophiocomina nigra (A), Crossaster papposus (F), Asterias rubens (P), Luidia ciliaris (P), Porania pulvillus (F), small teleost spp. (F).	SS.SMx.CMx.OphMx	-	-
D6-V04.11	Sand (73%) with shell and stone gravel (15%), pebbles (10%), shells (2%).	Stones sparsely encrusted with serpulid worms (P) and supporting hydroids (R), Omalosecosa ramulosa (R) and filamentous/filiform red algae (R). Cerianthus lloydii (C), Munida rugosa (P), Cancer pagurus (P), Pecten maximus (P), Porania pulvillus (F), Ophiocomina nigra (R), teleost sp. (P).	SS.SMx.CMx.ClloMx	-	-
D6-V04.12	Muddy sand (90%) with shell gravel (9%) and shells (1%).	Shells support sparse hydroids (R) and Omalosecosa ramulosa (R). Sediment with megafaunal burrow, possibly that of Nephrops norvegicus, and worm casts. Cerianthus lloydii (R), Paguridae sp. (R), Buccinum undatum (R), Turritella communis shells (P, including probably some occupied). Munida rugosa (R), Pecten maximus (R), Cancer pagurus (P), Liocarcinus spp.? (R), Luidia ciliaris (P), Porania pulvillus (O), Ophiura ophiura (R), teleost spp. (R) including Callionymus sp.?	SS.SSa.CMuSa	-	-
T4-V04	Soft mud.	Mud densely burrowed, particularly by Calocaris macandreae and Nephrops norvegicus (C, 59 animals seen). Calocaris density is difficult to determine due to operlapping clusters of burrows and burrows of other species but is at least C overall and A at least locally. Virgularia mirabilis (R), Cerianthus lloydii (R), Urticina sp.? (R), Caridea sp. (R), Buccinum undatum (R), Crossaster papposus (P), Asterias rubens (O), teleost spp. (R), Pleuronectiformes sp. (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T4-V05	Soft mud.	Mud densely burrowed, particularly by Calocaris macandreae and Nephrops norvegicus (C, 63 animals seen). Calocaris density is difficult to determine due to operlapping clusters of burrows and burrows of other species but is at least C overall and A at least locally. Cerianthus lloydii (R), Sabella pavonina (R), Caridea sp. (O), Neptunea antiqua? (R), Asterias rubens (O), teleost spp. (R), Pleuronectiformes sp. (O). Possible trawl scars visible intermittently (e.g. 00:11:15 - 00:11:50, 00:18:59 - 00:19:09, 01:03:16 - 01:04:12).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T2-V02	Soft mud.	Moderate density of megafaunal burrows including Nephrops norvegicus (C, 175 animals seen) and smaller burrows including Calocaris macandreae (P). Virgularia mirabilis (R), Asterias rubens (R), teleost spp. (O).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T2-V03	Soft mud.	Moderate density of megafaunal burrows including Nephrops norvegicus (C, 186 animals seen) and smaller burrows including Calocaris macandreae (P). Virgularia mirabilis (R), Pennatula phosphorea (R), Liocarcinus spp. (R), teleost spp. (O) including Pleuronectiformes spp. (P).	SS.SMu.CFiMu.SpnMeg	-	BM:SB
T2-V04	Cohesive muddy sand (99%) with shell gravel (1%) and shells (<1%), particularly Arctica islandica.	Sediment with worm casts, emergent infaunal tubes and sparse megafaunal burrows, mostly small but some larger including possibly Nephrops norvegicus (O). Munida rugosa (F), Pagurus bernhardus (R), Liocarcinus spp. (R), Cancer pagurus (O), Turritella communis (P), Buccinum undatum (R), Aequipecten opercularis (R), Asterias rubens (O), Luidia ciliaris (P), Porania pulvillus (O, locally F), Astropecten irregularis? (R), teleost spp. (R), Callionymus spp. (R), Chelidonichthys lucerna (P). Fishing debris (rubber glove).	SS.SMu.CSaMu	-	-
T2-V05	Cohesive muddy sand with shell gravel (<1%) and shells (<1%), particularly Arctica islandica.	Sediment with worm casts, bivalve siphons (P), small mounds, emergent infaunal tubes and low density of megafaunal burrows including Nephrops norvegicus (F, 4 animals seen); fairly large areas with few or no burrows. Hydroids (R), Munida rugosa (F), Pagurus bernhardus (R), Paguridae sp. (P in Turritella shell), Liocarcinus spp. (R), Cancer pagurus (O), Turritella communis (shells present and some probably occupied), Asterias rubens (O), Luidia ciliaris (O), Porania pulvillus (R), Ophiocomina nigra (R, locally C), teleost spp. (R). Habitat intermediate in nature between CSaMu and SpnMeg.	SS.SMu.CSaMu	-	-
T2-V06	Cohesive muddy sand or possibly sandy mud with sparse shell gravel (<1%) and shells (<1%).	Poor visibility. Sediment with worm casts and faunal tracks and lightly burrowed by megafaunal species including Nephrops norvegicus (C, 10 animals seen) and probably Calocaris macandreae (P) and supporting Pennatula phosphorea (R). Munida rugosa (O), Liocarcinus spp. (R), Cancer pagurus (P), Turritella communis (shells present and some possibly occupied), Aequipecten opercularis? (R), Asterias rubens (O), Porania pulvillus (R), Ophiocomina nigra (R, locally C), Ophiura ophiura (P), teleost spp. (R), Callionymus sp. (R). Habitat considered a poorly developed example of SpnMeg. Sediment with parallel streaks visible at several points along run (00:05:36 - 00:08:17, 00:14:00 - 00:14:56, 00:42:02 - 00:42:44), presumably trawl scars.	SS.SMu.CFiMu.SpnMeg	-	BM:SB
D6-V05.01	Mixed sediment of silty sand (55%) and gravel (40%, much higher locally), with pebbles (2%) and shells (3%).	Stones and shells sparsely encrusted with serpulid worms (P) and supporting hydroids (R), Urticina spp. (R), Flustra foliacea (R), tubular bryozoan (R), solitary ascidians (R) and filamentous/filiform red algae (O). Munida rugosa (F), Caridea sp. (P), Pagurus bernhardus (R), Liocarcinus spp. (R), Cancer pagurus (P), Aequipecten opercularis (R), Porania pulvillus (F), Astropecten irregularis (R), Luidia ciliaris (O), teleost sp. (P), Callionymus spp. (R), sparse emergent infaunal tubes.	SS.SMx.CMx	-	-
D6-V05.02	Silty fine sand (95%), with shell gravel (4%) and shells (1%).	Sediment with small holes and worm casts. Shell material encrusted with serpulid worms (R) and supporting hydroids (R), filamentous/filiform red algae (R), Flustra foliacea (R) and Urticina spp. (R). Munida rugosa (R), Pagurus bernhardus (R), Liocarcinus spp. (R), Cancer pagurus (P), Turritella communis (P), Buccinum undatum (R), Pecten maximus (R), Aequipecten opercularis (R), Porania pulvillus (O), Luidia ciliaris (O), Astropecten irregularis (R), Callionymus spp. (R).	SS.SSa.CMuSa	-	-
D6-V05.03	Mixed sediment of silty sand (60%) and shell gravel (25%), with pebbles (5%), shells (4%), cobbles (5%) and boulders (1%).	Stones and shells sparsely encrusted with serpulid worms (P) and supporting hydroids (R), Securiflustra securifrons? (R) and solitary ascidians (R). Munida rugosa (F), Paguridae sp. (P), Pecten maximus (P), Marthasterias glacialis (P), Porania pulvillus (F), Echinus esculentus (P), teleost sp. (P), sparse emergent infaunal tubes.	SS.SMx.CMx	-	-
D6-V05.04	Silty, fine sand (95%) with shell gravel (4%) and shells (1%); cobbles (<1%).	Shell material supporting sparse hydroids (R). Liocarcinus sp. (R), Turritella communis shells (P), Marthasterias glacialis (P), Porania pulvillus (O), teleost sp. (P).	SS.SSa.CMuSa	-	-
D4-V04.01	Mixed substrate of silty sand (35%) with gravel (10%) and pebbles (10%); areas of silted cobbles (30%) and boulders (15%).	Stones encrusted with serpulid worms (R) and supporting patchy hydroids (O) and Omalosecosa ramulosa (R). Asterias rubens (C). Considered a mosaic of BrAs in boulder/cobble areas and CMx elsewhere. Poor visibility.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V04.02	Silty, fine sand (85%) with shell gravel (10%) and shells (5%).	Sediment with small mounds, worm casts, emergent infaunal tubes and Cerianthus lloydii? (R). Shells supporting sparse hydroid clumps (R). Cancer pagurus (P), Liocarcinus spp. (R), Pecten maximus (R), Asterias rubens (O), Luidia ciliaris (P), Astropecten irregularis (R), Porania pulvillus (O).	SS.SSa.CMuSa	-	-
D4-V04.03	Mixed substrate of silty sand (35%) and gravel (10%) with pebbles (20%) and shells (<1%); silted cobbles (25%) and boulders (10%), both denser locally.	Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting patchy turf of hydroids (O, locally F), as well as Ascidia mentula (P). Munida rugosa (O), Pecten maximus (O), Asterias rubens (F), Porania pulvillus (O), Marthasterias glacialis (C), Luidia ciliaris (F), teleost spp. (O). Sediment with small holes and mounds. Considered a mosaic of BrAs in dense boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V04.04	Shelly (10%), possibly silty sand with scattered shells (2%) and sparse boulders (<1%).	Sediment with small mounds. Cancer pagurus (P), Pecten maximus (R), Marthasterias glacialis (P), Asterias rubens (P), Astropecten irregularis (R), Porania pulvillus (O).	SS.SSa.CMuSa	-	-
D4-V04.05	Mixed substrate of sand (20%) and gravel (10%) with pebbles (25%) and shells (<1%); silted cobbles (30%) and boulders (15%), both denser locally.	Stones encrusted with pink coralline algae (R), Parasmittina trispinosa (R) and serpulid worms (P) and supporting patchy turf of hydroids (O, locally F), as well as Omalosecosa ramulosa (R) and Ascidia mentula (F). Munida rugosa (O), Galathea sp.? (P), Asterias rubens (C), Porania pulvillus (O), Marthasterias glacialis (F), Stichastrella rosea (P), Henricia sp. (P), Luidia ciliaris (P), Echinus esculentus (F), teleost spp. (O). Sediment with small mounds. Considered a mosaic of BrAs in dense boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V04.06	Megaripples of medium-coarse sand (75%) with gravel (25%), much of it concentrated in troughs; boulders (<1%).	Sparse visible life. Asterias rubens (P), Luidia ciliaris (P).	SS.SCS.CCS	-	-
D4-V04.07	Mixed substrate of sand (30%) and gravel (10%) with pebbles (20%); silted cobbles (25%) and boulders (15%), both denser locally.	Stones encrusted with pink coralline algae (R), Parasmittina trispinosa (R) and serpulid worms (P) and supporting patchy turf of hydroids (O, locally F) and Ascidia mentula (O). Munida rugosa (O), Calliostoma zizyphinum (P), Crosssaster papposus (F), Asterias rubens (C), Porania pulvillus (F), Marthasterias glacialis (P), Stichastrella rosea (P), Luidia ciliaris (F), Echinus esculentus (F), teleost spp. (O), Labrus mixtus (P), Callionymus sp. (O). Considered a mosaic of BrAs in dense boulder/cobble areas and CMx elsewhere.	SS.SMx.CMx, CR.LCR.BrAs	RF:ST	-
D4-V04.08	Mixed sediment of sand (55%) with gravel (35%), pebbles (5%), cobbles (3%), shells (2%) and boulders (<1%).	Stones encrusted with pink coralline algae (R) and serpulid worms (P) and supporting patchy turf of hydroids (R), Omalosecosa ramosa (R) and Ascidia mentula (R). Munida rugosa (O), Cancer pagurus (P), Pecten maximus (O), Asterias rubens (P), Porania pulvillus (O), Marthasterias glacialis (F), teleost spp. (O). Sediment with emergent infaunal tubes (P) and some small mounds.	SS.SMx.CMx	-	-
D4-V05.01	Cohesive muddy sand or sandy mud (98%) with shell gravel (1%), shells (1%) and sparse cobbles (<1%) and boulders (<1%).	Sediment with small holes, mounds, worm casts, emergent infaunal tubes and sparse megafaunal burrows. Shells and stones support hydroid clumps (R). Munida rugosa (R), Pagurus bernhardus (R), Liocarcinus spp. (O), Turritella communis (F), Pecten maximus (R), Antedon sp. (R), Asterias rubens (P), Luidia ciliaris (P), Porania pulvillus (O), Ophiura ophiura (locally C), Callionymus sp. (R), teleost spp. indet. (R).	SS.SMu.CSaMu	-	-
D4-V05.02	Muddy sand (60%) with gravel (10%), pebbles (20%), shells (2%) and cobbles (8%).	Stones supporting hydroids (O). Asterias rubens (P), Porania pulvillus (F), teleost sp. (P).	SS.SMx.CMx	-	-
D4-V05.03	Muddy sand (94%) with shell gravel (5%), pebbles (<1%) and shells (1%).	Hydroids (R), Liocarcinus spp. (O), Turritella communis shells (P), Porania pulvillus (O), very sparse burrows and small mounds. Poor visibility.	SS.SMu.CSaMu	-	-
D4-V05.04	Muddy sand (65%) with gravel (10%), pebbles (20%), cobbles (5%), shells (<1%) and boulders (<1%).	Stones supporting patchy hydroids (O) and Omalosecosa ramosa (R). Munida rugosa (F), Pecten maximus (O), Porania pulvillus (O), Marthasterias glacialis (P), Astropecten irregularis (O), teleost sp. (P), Callionymus sp. (P).	SS.SMx.CMx	-	-
D4-V05.05	Muddy sand (95%) with shell gravel (5%), pebbles (<1%), cobbles (<1%) and shells (<1%).	Hydroids (R), Munida rugosa (P), Turritella communis (P), Marthasterias glacialis (P), Porania pulvillus (P), Asterias rubens (P), Luidia ciliaris (P), teleost sp. (P), infaunal casts (P). Poor visibility.	SS.SMu.CSaMu	-	-
D4-V05.06	Mixed sediment of muddy sand (50%) with gravel (10%), silted pebbles (30%), cobbles (10%) and boulders (<1%).	Stones with hydroid turf (F). Pecten maximus (P), Marthasterias glacialis (P), Poor visibility.	SS.SMx.CMx	-	-
D4-V05.07	Muddy sand (95%) with shell gravel (5%), shells (<1%) and cobbles (<1%).	Stones with hydroids (R). Munida rugosa (P), Pecten maximus (O).	SS.SMu.CSaMu	-	-
D4-V05.08	Muddy sand (80%) with gravel (5%), pebbles (10%), cobbles (5%) and shells (<1%).	Stones with hydroids (O) and Omalosecosa ramulosa (R). Munida rugosa (P), Buccinum undatum eggs? (P), Pecten maximus (O).	SS.SMx.CMx	-	-
D4-V05.09	Muddy sand or sandy mud (98%), shell gravel (2%), shells (<1%) and cobbles (<1%).	Sediment with sparse small burrows and mounds. Hydroids (R), Pagurus bernhardus? (P), Liocarcinus spp. (O), Turritella communis (P), Porania pulvillus (O), Luidia ciliaris (P).	SS.SMu.CSaMu	-	-

Annex 3: Biotopes recorded with sites of occurrence and illustrative photograph or video frame grab

Biotope codes in bold, italic font are Protected Features. Emboldened site codes indicate provenance of image. See Connor et al. (2004) for full biotope description.

CR.LCR Low energy circalittoral rock

T1-3.02, T1-7.01, T1-7.03, T1-7.05

Brachiopod and ascidian communities biotope

CR.LCR.BrAs Brachiopods and ascidians

D1-1.03, D4-V02.02, D4-V02.04, D4-V02.06, D4-V04.01, D4-V04.03, D4-V04.05, D4-V04.07, D5-2.09

Antedon spp., solitary ascidians and fine hydroids on sheltered circalittoral rock biotope

CR.LCR.BrAs.AntAsH Antedon spp., solitary ascidians and fine hydroids on sheltered circalittoral rock

D3-5.02, D4-V01.02, D4-V01.05, D4-V01.08, D4-V01.11, D5-2.04

SS.SCS.ICS Infralittoral coarse sediment

D1-4.01

SS.SCS.CCS Circalittoral coarse sediment

D4-V04.06, D6-V03.02, D6-V03.04

Neopentadactyla mixta in circalittoral shell gravel or coarse sand biotope

SS.SCS.CCS.Nmix Neopentadactyla mixta in circalittoral shell gravel or coarse sand

D1-2.03

Note specimen of Neopentadactyla mixta on right

SS.SSa.CMuSa Circalittoral muddy sand

D3-2.01, D3-2.03, D3-3.04, D3-5.04, D4-V01.01, D4-V01.03, D4-V01.06, D4-V01.10, D4-V01.13, D4-V04.02, D4-V04.04, D5-1.03, D5-2.02, D5-2.06, D5-3.03, D6-V01.01, D6-V04.12, D6-V05.02, D6-V05.04

SS.SMu.CSaMu. Circalittoral sandy mud

D1-6.02, D1-6.04, D3-3.05, D3-5.05, D3-5.07, D4-V03, D4-V05.01, D4-V05.03, D4-V05.05, D4-V05.07, D4-V05.09, D5-1.02, D5-2.01, D5-2.07, D5-3.04, D5-4.04, D5-5.03, T2-V04, T2-V05, T5-3.02

SS.SMu.CFiMu Circalittoral fine mud

T1-3.02, T1-7.01, T1-7.03

Seapens and burrowing megafauna in circalittoral fine mud biotope

SS.SMu.CFiMu.SpnMeg Seapens and burrowing megafauna in circalittoral fine mud

D3-3.06, D3-6, D5-1.01, D5-2.08, D5-2.09, D5-2.10, T1-1, T1-2, T1-3.01, T1-3.03, T1-4, T1-5, T1-6, T1-7.02, T1-7.04, T1-8, T1-9, T2-V01, T2-V02, T2-V03, T2-V06, T3-V01, T3-V02, T3-V03.01, T3-V03.03, T3-V03.05, T3-V04, T4-V01, T4-V02, T4-V03, T4-V04, T4-V05, T5-1, T5-2, T5-3.01

SS.SMx.CMx Circalittoral mixed sediment

D1-1.03, D1-6.01, D1-6.03, D3-3.03, D3-5.06, D4-V01.02, D4-V01.04, D4-V01.05, D4-V01.07, D4-V01.09, D4-V01.12, D4-V02.01, D4-V02.02, D4-V02.03, D4-V02.04, D4-V02.05, D4-V02.06, D4-V02.07, D4-V04.01, D4-V04.03, D4-V04.05, D4-V04.07, D4-V04.08, D4-V05.02, D4-V05.04, D4-V05.06, D4-V05.08, D5-2.04, D5-2.05, D5-3.02, D6-V05.01, D6-V05.03, T3-V03.02, T3-V03.04, T3-V03.06

Cerianthus lloydii and other burrowing anemones in circalittoral muddy mixed sediment biotope

SS.SMx.CMx.ClloMx Cerianthus lloydii and other burrowing anemones in circalittoral muddy mixed sediment

D1-1.02, D1-2.04, D1-3.02, D1-5.02, D3-3.02, D3-4, D3-7.02, D5-6.02, D6-V01.02, D6-V02, D6-V03.01, D6-V04.01, D6-V04.03, D6-V04.05, D6-V04.07, D6-V04.09, D6-V04.11

Cerianthus lloydii with Nemertesia spp.and other hydroids in circalittoral muddy mixed sediment biotope

SS.SMx.CMx.ClloMx.Nem Cerianthus lloydii with Nemertesia spp.and other hydroids in circalittoral muddy mixed sediment

D1-4.04, D3-2.02, D3-5.01, D3-5.02, D3-5.03, D5-1.04, D5-2.03, D5-3.01, D5-4.02, D5-4.03, D5-5.02

Ophiothrix fragilis and/or Ophiocomina nigra brittlestar beds on sublittoral mixed sediment biotope

SS.SMx.CMx.OphMx Ophiothrix fragilis and/or Ophiocomina nigra brittlestar beds on sublittoral mixed sediment

D1-1.04, D1-4.03, D6-V04.02, D6-V04.04, D6-V04.06, D6-V04.08, D6-V04.10

SS.SMp.Mrl Maerl beds

D6-V03.03

Phymatolithon calcareum maerl beds with red seaweeds in shallow infralittoral clean gravel or coarse sand biotope

SS.SMp.Mrl.Pcal.R Phymatolithon calcareum maerl beds with red seaweeds in shallow infralittoral clean gravel or coarse sand

LBN-1.03, LBN-1.05

Kelp and seaweed communities on sublittoral sediment biotope

SS.SMp.KSwSS Kelp and seaweed communities on sublittoral sediment

D6-V03.05

Saccharina latissima and red seaweeds on infralittoral sediments biotope

SS.SMp.KSwSS.LsacR Saccharina latissima and red seaweeds on infralittoral sediments

LBN-1.02, LBN-1.04, LBN-1.06

Saccharina latissima and robust red algae on infralittoral gravel and pebbles biotope

SS.SMp.KSwSS.LsacR.Gv Saccharina latissima and robust red algae on infralittoral gravel and pebbles

D1-1.01, D1-2.01, D1-2.02, D1-3.01, D1-4.02, D1-5.01, D3-1, D3-3.01, D3-7.01, D5-4.01, D5-5.01, D5-6.01, LBN-1.01

Saccharina latissima and filamentous red algae on infralittoral sand biotope

SS.SMp.KSwSS.LsacR.Sa Saccharina latissima and filamentous red algae on infralittoral sand

D3-1

Annex 4: Priority Marine Feature (PMF), Protected Feature (PF) and qualifying Annex I habitat type abbreviations used in tables

PMFs & PFs

AI	Ocean quahog (Arctica islandica)
BM:SB	Burrowed mud: sea pens and burrowing megafauna
BM:FQ	Burrowed mud: Funiculina quadrangularis
KS	Kelp and seaweed communities on sublittoral sediment
MB	Maerl beds
MC	Maerl or coarse shell gravel with burrowing sea cucumbers
SE	Sandeels (Ammodytes marinus and A. tobianus)

Annex 1 habitats

RF:BR	Reef: bedrock
RF:ST	Reef: stony
SB:GS	Sandbanks slightly covered by seawater all the time: gravelly and clean sands
SB:KS	Sandbanks slightly covered by seawater all the time: kelp and seaweed
SB:MX	Sandbanks slightly covered by seawater all the time: mixed sediment
SB:MB	Sandbanks slightly covered by seawater all the time: maerl beds

NatureScot Research Report 1265 - Biological analyses of underwater video footage collected from around the Isle of Arran (Firth of Clyde) in 2015

Keywords

Background

Main findings

Acknowledgements

Introduction

Methods

Results

Seol Mara July 2015 survey

Box D3 (Figure 3)

Box T1 (Figure 4)

Box D5 and run LBN-1 (Figure 5)

Box D1 and T5 runs (Figure 6)

Alba na Mara September 2015 survey

Box T4 (Figure 7)

Box D6 (Figure 8)

Box T2 (Figure 9)

Box D4 (Figure 10)

Box T3 (Figure 11)

Discussion

References

Annex 1: Positional, temporal and depth details of video sequences recorded during the surveys

Annex 2: Physical and biological descriptions of the survey sites, with indicators of conservation importance of habitats and species

Annex 3: Biotopes recorded with sites of occurrence and illustrative photograph or video frame grab

Annex 4: Priority Marine Feature (PMF), Protected Feature (PF) and qualifying Annex I habitat type abbreviations used in tables

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